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Classification System: APG IV

Superregnum: Eukaryota
Regnum: Plantae
Cladus: Angiosperms
Cladus: Eudicots
Cladus: Core eudicots
Cladus: Asterids
Cladus: Campanulids
Ordo: Asterales

Familia: Asteraceae
Subfamilia: Asteroideae
Tribus: Astereae
Subtribus: Chrysopsidinae
Genus: Heterotheca
Species: H. arizonica – H. brandegei – H. camporum – H. canescens – H. fastigiata – H. fulcrata – H. grandiflora – H. gypsophila – H. inuloides – H. jonesii – H. leptoglossa – H. marginata – H. mexicana – H. monarchensis – H. mucronata – H. oregona – H. pumila – H. rutteri – H. sessiliflora – H. shevockii – H. stenophylla – H. subaxillaris – H. villosa – H. viscida – H. zionensis
Name

Heterotheca Cass.

Type species: Heterotheca lamarckii Cass.

Synonyms

Diplocoma D.Don ex D.Don
Chrysopsis subg. Ammodia (Nutt.) A.Gray
Ammodia Nutt.
Heterotheca sect. Ammodia (Nutt.) V.L.Harms
Calycium Elliott

References

Cass., Bulletin des Sciences, par la Societe Philomatique 1817: 137. 1817.

Links

International Plant Names Index. 2017. Heterotheca. Published online. Accessed: Dec. 10 2017.
The Plant List 2013. Heterotheca in The Plant List Version 1.1. Published on the internet. Accessed: 2017 Dec. 10.
Tropicos.org 2017. Heterotheca. Missouri Botanical Garden. Published on the internet. Accessed: 2017 Dec. 10.
Hassler, M. 2017. Heterotheca. World Plants: Synonymic Checklists of the Vascular Plants of the World In: Roskovh, Y., Abucay, L., Orrell, T., Nicolson, D., Bailly, N., Kirk, P., Bourgoin, T., DeWalt, R.E., Decock, W., De Wever, A., Nieukerken, E. van, Zarucchi, J. & Penev, L., eds. 2017. Species 2000 & ITIS Catalogue of Life. Published on the internet. Accessed: 2017 Dec 10. Reference page.

Vernacular names
English: Goldenaster, camphorweed, telegraph weed
русский: Гетеротека

Heterotheca, (common names goldenasters, camphorweed, or telegraph weed)[5] are North American plants in the sunflower family.[2][6]

Etymology

Heterotheca comes from Ancient Greek ἕτερος héteros "other, different" and θήκη thḗkē "case, chest", and refers to the fact that, in some species in the genus, the cypselae (achenes containing seed) of the disk and ray florets have different shapes.[5][7]
Description, biology

These are annual and perennial herbs bearing daisy-like flower heads with yellow disc florets and usually yellow ray florets, associated with mesic to xeric habitats across North America. Several species now included in Heterotheca were previously classified in the genus Chrysopsis[1][5]

Heterotheca species are often used as food plants by the larvae of some Lepidoptera species including Schinia lynx, Schinia nubila and Schinia saturata (all of which have been recorded on Heterotheca subaxillaris).
Chemistry

The leaf volatiles from which the name "camphorweed" is derived include camphor, but as a minor constituent (less than 2%); of the 41 documented volatiles in a study of Heterotheca subaxillaris, for example, caryophyllene, pinene, borneol, myrcene, and limonene each comprised over 5% of the total.[8]
Species

Heterotheca barbata (Rydb.) Semple - Spokane golden aster - Washington Idaho
Heterotheca brandegeei (B.L.Rob. & Greenm.) Semple - northern Baja California
Heterotheca camporum (Greene) Shinners - Arkansas Missouri Iowa Illinois Ohio Michigan Kentucky Tennessee Mississippi Alabama Georgia Virginia North Carolina New Jersey
Heterotheca canescens (DC.) Shinners - Nuevo León, Texas New Mexico Oklahoma Colorado Kansas Missouri Iowa
Heterotheca fastigiata (Greene) V.L.Harms - California
Heterotheca fulcrata (Greene) Shinners - Coahuila, Tamaulipas, Arizona New Mexico Texas Nevada Utah Colorado Wyoming Idaho
Heterotheca grandiflora - Telegraphweed - Baja California, California Arizona Nevada Utah
Heterotheca gypsophila B.L.Turner - Nuevo León
Heterotheca inuloides Cass. - Mexican arnica - from Nuevo León to Oaxaca
Heterotheca jonesii (S.F.Blake) S.L.Welsh & N.D.Atwood - Utah
Heterotheca leptoglossa DC. - Guanajuato, Chihuahua, Sonora, Sinaloa, Jalisco, San Luis Potosí
Heterotheca marginata Semple - Arizona
Heterotheca mexicana V.L.Harms ex B.L.Turner - Durango
Heterotheca monarchensis D.A.York, Shevock & Semple - monarch golden aster - Fresno County in California
Heterotheca mucronata V.L.Harms ex B.L.Turner - Nuevo León, Coahuila, Tamaulipas
Heterotheca oregona (Nutt.) Shinners
Heterotheca pumila (Greene) Semple - Colorado Wyoming Utah New Mexico
Heterotheca rutteri (Rothr.) Shinners - Sonora, Arizona
Heterotheca sessiliflora (Nutt.) Shinners - Sessileflower false golden aster - Baja California, California
Heterotheca shevockii (Semple) Semple - Kern Canyon false golden aster - Kern County in California
Heterotheca stenophylla (A.Gray) Shinners - Texas New Mexico Oklahoma Colorado Wyoming Kansas Nebraska South Dakota Iowa Minnesota
Heterotheca subaxillaris (Lam.) Britton & Rusby - Camphorweed, camphor weed, false goldenaster[7] - widespread from Belize to California, South Dakota, + Massachusetts
Heterotheca villosa (Pursh) Shinners
Heterotheca viscida (A.Gray) V.L.Harms - Arizona New Mexico Texas
Heterotheca zionensis Semple - Zion golden aster - Arizona New Mexico Texas Utah Idaho Wyoming

Sources:[1][9]
Formerly included

Many species have been included in Heterotheca at various times in the past, but now regarded as more suitable for other genera. The most common of these is Chrysopsis, but others include Aster Bradburia Osbertia Munnozia Pityopsis + Tomentaurum.[1]
References

Flann, C (ed) 2009+ Global Compositae Checklist
"Heterotheca". Tropicos. Missouri Botanical Garden.
"Heterotheca". The Plant List. Missouri Botanical Garden. Royal Botanic Gardens, Kew.
The Plant List, search for Heterotheca
Semple, John C. (2006). "Heterotheca". In Flora of North America Editorial Committee (ed.). Flora of North America North of Mexico (FNA). 20. New York and Oxford. Retrieved 16 July 2016 – via eFloras.org, Missouri Botanical Garden, St. Louis, MO & Harvard University Herbaria, Cambridge, MA.
Cassini, Alexandre Henri Gabriel de. 1817. Bulletin des Sciences, par la Societe Philomatique 1817: 137
C. Williamson, Gerald (2016). "US Wildflower - Camphorweed, Camphor Weed, False Goldenaster - Heterotheca subaxillaris". US Wildflowers. Retrieved 16 July 2016.
Lincoln, D.E., B.M. Lawrence. 1984. The volatile constituents of camphorweed, Heterotheca subaxillaris. Phytochemistry 23(4):933-934
"Heterotheca". County-level distribution maps from the North American Plant Atlas (NAPA). Biota of North America Program (BONAP). 2014.

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