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Classification System: APG IV

Superregnum: Eukaryota
Regnum: Plantae
Cladus: Angiosperms
Cladus: Eudicots
Cladus: Core eudicots
Cladus: Asterids
Cladus: Lamiids
Ordo: Gentianales

Familia: Rubiaceae
Subfamilia: Rubioideae
Tribus: Knoxieae
Genera: BatopedinaCarphaleaChamaepentas – Dirichletia – Dolichopentas – Knoxia – Lathraeocarpa – Otiophora – Otomeria – Paracarphalea – Paraknoxia – Parapentas – Pentanisia – Pentas – Phyllopentas – Rhodopentas – Triainolepis

Knoxieae Benth. & Hook.f., Gen. Pl. 2: 9, 21. 1873.
Type genus: Knoxia L.


Knoxiinae K.Schum. in Engl. & Prantl, Nat. Pflanzenfam. IV, 4: 16, 89. 1891.
Knoxioideae Luerss., Handb. Syst. Bot. 2: 1083. 1882.
Type genus: Knoxia L.


Bentham, G. & Hooker, J.D. 1873. Genera Plantarum ad exemplaria imprimis in herbariis Kewensibus 2: 9, 21.
Ferm, J., Kårehed, J.E.S.P.E.R., Bremer, B. & Razafimandimbison, S.G. 2016. Paracarphalea, a new genus of the coffee family segregated from the Malagasy endemic genus Carphalea (Rubiaceae, Rubioideae, Knoxieae). Phytotaxa 263(2): 98-112. DOI: 10.11646/phytotaxa.263.2.2 PDF from ResearchGate Reference page.
Kårehed, J. & Bremer, B. 2007. The systematics of Knoxieae (Rubiaceae) — molecular data and their taxonomic consequences. Taxon 56(4): 1051–1076. DOI: 10.2307/25065904 JSTOR PDF. Reference page.

Knoxieae is a tribe of flowering plants in the family Rubiaceae and contains about 131 species in 16 genera. Its representatives are found in Tropical and Southern Africa, the islands in the Western Indian Ocean, the Arabian Peninsula, Tropical and Subtropical Asia, and Northern Australia.[1]


Currently accepted names[1][2][3][4]

Batopedina Verdc. (3 sp)
Carphalea Juss. (3 sp)
Chamaepentas Bremek. (6 sp)
Dirichletia Klotzsch (5 sp)
Dolichopentas Kårehed & B.Bremer (4 sp)
Knoxia L. (13 sp)
Otiophora Zucc. (18 sp)
Otomeria Benth. (8 sp)
Paracarphalea Razafimandimbison, Ferm, B.Bremer & Kårehed (3 sp)
Paraknoxia Bremek. (1 sp)
Parapentas Bremek. (3 sp)
Pentanisia Harv. (19 sp)
Pentas Benth. (16 sp)
Phyllopentas (Verdc.) Kårehed & B.Bremer (14 sp)
Rhodopentas Kårehed & B.Bremer (2 sp)
Triainolepis Hook.f. (13 sp)


Afroknoxia Verdc. = Knoxia
Baumannia K.Schum. = Knoxia
Calanda K.Schum. = Pentanisia
Chlorochorion Puff & Robbr. = Pentanisia
Cuncea Buch.-Ham. ex D.Don = Knoxia
Dentillaria Kuntze = Knoxia
Diotocarpus Hochst. = Pentanisia
Holocarpa Baker = Pentanisia
Mericocalyx Bamps = Otiophora
Neobaumannia Hutch. & Dalziel = Knoxia
Neopentanisia Verdc. = Pentanisia
Neurocarpaea R.Br. ex Hiern = Pentas
Neurocarpaea R.Br. = Pentas
Orthostemma Wall. ex Voigt = Pentas
Otocephalus Chiov. = Pentanisia
Paratriaina Bremek. = Triainolepis
Pentacarpaea Hiern = Pentanisia
Pentacarpus Post & Kuntze = Pentanisia
Placopoda Balf.f. = Dirichletia
Princea Dubard & Dop = Triainolepis
Tapinopentas Bremek. = Otomeria
Thyridocalyx Bremek. = Triainolepis
Vignaldia A.Rich. = Pentas
Vissadali Adans. = Knoxia


"World Checklist of Rubiaceae". Retrieved 20 April 2016.
Robbrecht E, Manen J-F (2006). "The major evolutionary lineages of the coffee family (Rubiaceae, angiosperms). Combined analysis (nDNA and cpDNA) to infer the position of Coptosapelta and Luculia, and supertree construction based on rbcL, rps16, trnL-trnF and atpB-rbcL data. A new classification in two subfamilies, Cinchonoideae and Rubioideae". Systematic Geography of Plants. 76: 85–146.
Bremer B (2009). "A review of molecular phylogenetic studies of Rubiaceae". Annals of the Missouri Botanical Garden. 96: 4–26. doi:10.3417/2006197.
Bremer B, Eriksson E (2009). "Time tree of Rubiaceae: phylogeny and dating the family, subfamilies, and tribes". International Journal of Plant Sciences. 170: 766–793. doi:10.1086/599077.

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