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Classification System: APG IV

Superregnum: Eukaryota
Regnum: Plantae
Cladus: Angiosperms
Cladus: Eudicots
Cladus: Core eudicots
Cladus: Asterids
Cladus: Lamiids
Ordo: Gentianales

Familia: Rubiaceae
Subfamilia: Ixoroideae
Tribus: Gardenieae
Genus: Tocoyena
Species: T. arenicola – T. brasiliensis – T. brevifolia – T. bullata – T. costanensis – T. cubensis – T. foetida – T. formosa – T. guianensis – T. hispidula – T. longiflora – T. megistantha – T. neglecta – T. orinocensis – T. pendulina – T. pittieri – T. sellowiana – T. tomentosa – T. uniflora – T. viscidula – T. williamsii

Tocoyena Aubl., Hist. Pl. Guiane 1: 131 (1775)

Type species: Tocoyena longiflora Aubl., Hist. Pl. Guiane: 131. (1775)


Ucriana Willd., Sp. Pl. 1: 961. 1798.


Aublet, J.B.C.F. 1775. Histoire des Plantes de la Guiane Françoise 1: 131.
Govaerts, R. et al. 2019. Tocoyena in World Checklist of Selected Plant Families. The Board of Trustees of the Royal Botanic Gardens, Kew. Published on the internet. Accessed: 2019 Apr. 27. Reference page.
International Plant Names Index. 2019. Tocoyena. Published online. Accessed: Apr. 27 2019. 2019. Tocoyena. Missouri Botanical Garden. Published on the internet. Accessed: 2019 Apr. 27.

Tocoyena is a genus of plant in family Rubiaceae.[1]

Tocoyena contains the following species (incomplete list):[2][3][4]

Tocoyena atlantica R. Borges & Gaem
Tocoyena brasiliensis Mart.
Tocoyena bullata (Vell.) Mart.
Tocoyena foetida Poepp. & Endl.
Tocoyena formosa (Cham. & Schltdl.) K. Schum.
Tocoyena guyanensis K. Schum.
Tocoyena pittieri (Standl.) Standl.
Tocoyena selloana Schum.
Tocoyena sellowiana (Cham. & Schltdl.) K. Schum.


REVISION OF TOCOYENA (RUBIACEAE: GARDENIEAE) FROM THE STATES OF GOIÁS AND TOCANTINS AND A NEW SPECIES ENDEMIC TO WHITE-SAND AREAS IN THE BRAZILIAN CERRADO. Piero G. Delprete, Journal of the Botanical Research Institute of Texas, 9 December 2008, Vol. 2, No. 2, pages 983-993 (article)
DELPRETE, PIERO G. (2018-12-10). "Two new species of the tribe Sipaneeae (Rubiaceae) from white-sand areas of Brazilian and Colombian Amazon". Phytotaxa. 382 (1): 125. doi:10.11646/phytotaxa.382.1.7. ISSN 1179-3163.
Borges, Rodrigo L.; Gaem, Paulo Henrique; Roque, Nádia (2020-11-05). "A new species of Tocoyena (Rubiaceae, Gardenieae) from the Brazilian Atlantic Forest". Phytotaxa. 470 (3): 243–248. doi:10.11646/phytotaxa.470.3.4. ISSN 1179-3163.
de Souza, Renata Gomes; Goto, Bruno Tomio; Alves da Silva, Danielle Karla; Barbosa da Silva, Fábio Sérgio; Sampaio, Everardo V.S.B.; Maia, Leonor Costa (2010). "The role of arbuscular mycorrhizal fungi and cattle manure in the establishment of Tocoyena selloana Schum. in mined dune areas". European Journal of Soil Biology. 46 (3–4): 237–242. doi:10.1016/j.ejsobi.2010.04.004. ISSN 1164-5563.

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