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Classification System: APG IV

Superregnum: Eukaryota
Regnum: Plantae
Cladus: Angiosperms
Cladus: Eudicots
Cladus: Core eudicots
Cladus: Rosids
Cladus: Eurosids I
Ordo: Zygophyllales

Familia: Zygophyllaceae
Subfamiliae: Larreoideae - Morkillioideae - Seetzenioideae - Tribuloideae - Zygophylloideae
Genera: AugeaBalanites – Larreoideae – Fagonia – Gonopterodendron – Guaiacum – Kallstroemia – Kelleronia – Larrea – Melocarpum – Metharme – MiltianthusMorkillia – Neoluederitzia – Pintoa – Plectrocarpa – PorlieriaRoeperaSeetzeniaSericodesSisynditeTetraenaTribulopisTribulusViscainoaZygophyllum


Zygophyllaceae R.Br., Voy. Terra Austral. 2: 545 (1814), nom. cons.

Type genus: Zygophyllum L., Sp. Pl. 1: 385. (1753)


Zygophyllanae Doweld
Balanitaceae M.Roem., Fam. Nat. Syn. Monogr. 1: 26. 14. (1846) nom. cons.
Tribulaceae Trautv., Estestv. Istor. Gub. Kievsk. Uchebn. Okr. Bot. Sist.: 28. (1853)


Brown, R. 1814. A Voyage to Terra Australis 2: 545.
Sheahan, M.C. & Chase, M.W. 1996. A phylogenetic analysis of Zygophyllaceae R.Br. based on morphological, anatomical and rbcL DNA sequence data. Botanical Journal of the Linnean Society 122(4): 279–299. DOI: 10.1111/j.1095-8339.1996.tb02077.x Open access Reference page.
Sheahan, M.C. & Chase, M.W. 2000. Phylogenetic relationships within Zygophyllaceae based on DNA sequences of three plastid regions, with special emphasis on Zygophylloideae. Systematic Botany 25(2): 371–384. DOI: 10.2307/2666648 Paywall JSTOR Reference page.
Godoy-Bürki, A.C., Acosta, J.M. & Aagesen, L. 2018. Phylogenetic relationships within the New World subfamily Larreoideae (Zygophyllaceae) confirm polyphyly of the disjunct genus Bulnesia. Systematics and Biodiversity 16(5): 453-468. DOI: 10.1080/14772000.2018.1451406 Paywall ResearchGate Reference page.
Govaerts, R. et al. 2020. Zygophyllaceae in Kew Science Plants of the World online. The Board of Trustees of the Royal Botanic Gardens, Kew. Published on the internet. Accessed: 2020 Nov. 15. Reference page.
Hassler, M. 2020. Zygophyllaceae. World Plants: Synonymic Checklists of the Vascular Plants of the World In: Roskovh, Y., Abucay, L., Orrell, T., Nicolson, D., Bailly, N., Kirk, P., Bourgoin, T., DeWalt, R.E., Decock, W., De Wever, A., Nieukerken, E. van, Zarucchi, J. & Penev, L., eds. 2020. Species 2000 & ITIS Catalogue of Life. Published on the internet. Accessed: 2020 Sept. 7. Reference page.
International Plant Names Index. 2020. Zygophyllaceae. Published online. Accessed: Nov. 15 2020.
Stevens, P.F. 2001 onwards. Angiosperm Phylogeny Website. Version 14, July 2017 [and more or less continuously updated since]. Online. Reference page.

Vernacular names
Deutsch: Jochblattgewächse
English: Creosote Bush Family
español: Zigofiláceas
suomi: Mehikorvakasvit
한국어: 남가새과
македонски: Парнолисни
русский: Парнолистниковые
Türkçe: Yabani kimyongiller
українська: Парнолистникові
中文: 蒺藜科

The Zygophyllaceae is a family of flowering plants that contains the bean-caper and caltrop. It includes around 285 species in 22 genera.

In the APG III system of classification, the families Zygophyllaceae and Krameriaceae compose the order Zygophyllales. In the previous version of their classification system, the Angiosperm Phylogeny Group had included the option of placing Krameria within Zygophyllaceae.

Zygophyllaceae is divided into five subfamilies. A molecular phylogeny of the family was published in 2000. Phylogenies of groups within the family have also been published. A phylogeny of the southern African species of Zygophyllum was published in 2008.

Peganum was in Zygophyllaceae before being moved to the newly created family Nitrariaceae.


Subfamily Larreoideae

* Bulnesia Gay
* Guaiacum L.
* Larrea Cav.
* Pintoa Gay
* Porlieria Ruiz & Pav.[2]

Subfamily Morkillioideae

* Morkillia Rose & J.H.Painter

* Sericodes A.Gray
* Viscainoa Greene[3]

Subfamily Seetzenioideae

* Seetzenia R.Br. ex Decne.[4]

Subfamily Tribuloideae

* Balanites Delile
* Kallstroemia Scop.
* Kelleronia Schinz

* Neoluederitzia Schinz
* Sisyndite E.Mey. ex Sond.
* Tribulopis R.Br.
* Tribulus L.[5]

Subfamily Zygophylloideae

* Augea Thunb.
* Fagonia L.
* Halimiphyllum (Engl.) Boriss.

* Melocarpum (Engl.) Beier & Thulin
* Roepera A.Juss.
* Tetraena Maxim.
* Zygophyllum L.[6]


* Izozogia G.Navarro
* Metharme Phil. ex Engl.
* Plectrocarpa Gillies ex Hook. & Arn.[7]


1. ^ "Zygophyllaceae R. Br., nom. cons.". Germplasm Resources Information Network. United States Department of Agriculture. 2003-01-17. http://www.ars-grin.gov/cgi-bin/npgs/html/family.pl?1210. Retrieved 2009-09-25.
2. ^ "Genera of Zygophyllaceae subfam. Larreoideae". Germplasm Resources Information Network. United States Department of Agriculture. http://www.ars-grin.gov/cgi-bin/npgs/html/gnlist.pl?2051. Retrieved 2010-08-15.
3. ^ "Genera of Zygophyllaceae subfam. Morkillioideae". Germplasm Resources Information Network. United States Department of Agriculture. http://www.ars-grin.gov/cgi-bin/npgs/html/gnlist.pl?2054. Retrieved 2010-08-15.
4. ^ "Genera of Zygophyllaceae subfam. Seetzenioideae". Germplasm Resources Information Network. United States Department of Agriculture. http://www.ars-grin.gov/cgi-bin/npgs/html/gnlist.pl?2052. Retrieved 2010-08-15.
5. ^ "Genera of Zygophyllaceae subfam. Tribuloideae". Germplasm Resources Information Network. United States Department of Agriculture. http://www.ars-grin.gov/cgi-bin/npgs/html/gnlist.pl?2053. Retrieved 2010-08-15.
6. ^ "Genera of Zygophyllaceae subfam. Zygophylloideae". Germplasm Resources Information Network. United States Department of Agriculture. http://www.ars-grin.gov/cgi-bin/npgs/html/gnlist.pl?2050. Retrieved 2010-08-15.
7. ^ "Genera of Zygophyllaceae". Germplasm Resources Information Network. United States Department of Agriculture. http://www.ars-grin.gov/cgi-bin/npgs/html/gnlist.pl?1210. Retrieved 2010-08-15.

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