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Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Endopterygota
Superordo: Panorpida
Cladus: Amphiesmenoptera
Ordo: Lepidoptera
Subordo: Glossata
Cladus: Coelolepida
Cladus: Myoglossata
Cladus: Neolepidoptera
Infraordo: Heteroneura
Cladus: Eulepidoptera
Cladus: Ditrysia
Cladus: Apoditrysia
Cladus: Obtectomera
Superfamilia: Papilionoidea

Familia: Hesperiidae
Subfamilia: Pyrginae
Tribus: Carcharodini
Subtribus: Carcharodina
Genus: Agyllia
Species (3): A. agylla – A. asterodia – A. kituina

Agyllia Grishin in Zhang et al., 2020: 186.

Type species: Pyrgus agylla Trimen, 1889.

Gender: feminine

Zhang, J., Brockmann, E., Cong, Q., Shen, J.H. & Grishin, N.V. 2020. A genomic perspective on the taxonomy of the subtribe Carcharodina (Lepidoptera: Hesperiidae: Carcharodini). Zootaxa 4748(1): 182–194. DOI: 10.11646/zootaxa.4748.1.10. Paywall. Reference page.

Vernacular names
English: Sandmen

Agyllia is a genus of spread-wing skippers in the butterfly family Hesperiidae, erected in 2020. There are at least three described species in Agyllia, all found in Africa. These species were formerly members of the genus Spialia.[1][2]

These species belong to the genus Agyllia:

Agyllia agylla (Trimen, 1889)
Agyllia asterodia (Trimen, 1864)
Agyllia kituina (Karsch, 1896)


Zhang, Jing; Brockmann, Ernst; Cong, Qian; Shen, Jinhui; et al. (2020). "A genomic perspective on the taxonomy of the subtribe Carcharodina (Lepidoptera: Hesperiidae: Carcharodini)". Zootaxa. 4748 (1). doi:10.11646/zootaxa.4748.1.10.

Li, Wenlin; Cong, Qian; Shen, Jinhui; Zhang, Jing; et al. (2019). "Genomes of skipper butterflies reveal extensive convergence of wing patterns". Proceedings of the National Academy of Sciences. 116 (13). doi:10.1073/pnas.1821304116. PMC 6442542.

Further reading

Zhang, Jing; Cong, Qian; Shen, Jinhui; Opler, Paul A.; et al. (2021). "Genomics-guided refinement of butterfly taxonomy" (PDF). The Taxonomic Report of the International Lepidoptera Survey. 9 (3). ISSN 2643-4806.

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