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Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Endopterygota
Superordo: Hymenopterida
Ordo: Hymenoptera
Subordo: Apocrita
Superfamilia: Ichneumonoidea

Familia: Ichneumonidae
Subfamilia: Banchinae
Tribus: Atrophini - Banchini - Glyptini - incertae sedis
Overview of genera

Hapsinotus – Ichnoverruca – Lissonota – Pristiboea – Terrylee – Valdiviglypta

Banchinae Wesmael, 1845

Bennett, A.M.R. 2009: The Ichneumonidae (Hymenoptera) of Fiji: keys to subfamilies and genera with a review of the species of Anomaloninae, Banchinae, Brachycyrtinae and Diplazontinae. In: Evenhuis, N.L. & Bickel, D.J. (eds.), Fiji Arthropods XIV. Bishop Museum occasional papers, 105: 3–68. PDF
Broad, G.R.; Sääksjärvi, I.E.; Veijalainen, A.; Notton, D.G. 2011: Three new genera of Banchinae (Hymenoptera: Ichneumonidae) from Central and South America. Journal of natural history, 45(21&22): 1311–1329. DOI: 10.1080/00222933.2011.552809
Kusigemati, K. 1987: Some Ephialtinae, Banchinae, Porizontinae, Mesochorinae, Metopiinae and Acaenitinae of Formosa (Hymenoptera: Ichneumonidae). Memoirs of the Kagoshima University Research Center for the South Pacific, 8(1): 21–33.
Sheng, M-L., Sun, S-P., Wang, X-N. & Wu, H-W. 2018. A new genus and species of subfamily Banchinae (Hymenoptera, Ichneumonidae) from China. Zootaxa 4413(3): 541–550. DOI: 10.11646/zootaxa.4413.3.8 Paywall Reference page.

Banchinae is a subfamily of ichneumonid parasitoid wasps containing about 1,500 species; the genera Glypta and Lissonota are very large. The three tribes (Banchini, Glyptini and Atrophini) are all distributed worldwide.[1][2]

In older treatments, the Lycorininae, Neorhacodinae and Stilbopinae are often included in the Banchinae; newer works usually consider them separate families.[3]

All banchines are koinobiont endoparasites of Lepidoptera. The Glyptini parasitise Tortricoidea. Atrophini parasitise a wider range of small moths. Species of Lissonota have long ovipositors able to reach deep wood-boring Lepidoptera such as Cossidae.[4] Banchinae and Campopleginae are the only subfamilies of Ichneumonidae known to have polydnaviruses.[5]

Most Banchinae have a stalked diamond-shaped areolet. A lobe of the propodeum projects over the middle coxae. The propodeum has few ridges (carinae), and the face is described as goat-like.


These 47 genera belong to the subfamily Banchinae:[6][7][8]

Agathilla Westwood, 1882 c g b
Alloplasta Förster, 1869 c g
Amphirhachis Townes, 1969 c g
Apophua Morley, 1913 c g
Arenetra Holmgren, 1859 c g b
Atropha Kriechbaumer, 1894 c g
Australoglypta Gauld, 1977 c g
Banchopsis Rudow, 1886 c g
Banchus Fabricius, 1798 c g b
Brachychroa Townes & Townes, 1978 c g
Catadacus Townes, 1969 c g
Cecidopimpla Brèthes, 1920 c g
Cephaloglypta Obrtel, 1956 c g
Ceratogastra Ashmead, 1900 c g b
Cryptopimpla Taschenberg, 1863 c g b
Diradops Townes, 1946 c g b
Eudeleboea Blanchard, 1936 g
Exetastes Gravenhorst, 1829 c g b
Geraldus Fitton, 1987 c g
Glypta Gravenhorst, 1829 c g b
Glyptopimpla Morley, 1913 c g
Helotorus Townes, 1978 c g
Himertosoma Schmiedeknecht, 1900 c g
Isomeris Townes, 1969
Leptobatopsis Ashmead, 1900 c g
Levibasis Townes, 1969 c g
Lissonota Gravenhorst, 1829 c g b
Lissonotidea Hellen, 1949 c g
Loxodocus Townes, 1969 c g
Mnioes Townes, 1946 c g
Neoexetastes Graf, 1984 c g
Odinophora Förster, 1869 c g
Philogalleria Cameron, 1912 c g
Rynchobanchus Kriechbaumer, 1894 c g
Sachtlebenia Townes, 1963 c g
Sjostedtiella Szépligeti, 1908 c g
Sphelodon Townes, 1966 c g b
Spilopimpla Cameron, 1904 c g
Stilbops Förster, 1869 c g
Syzeuctus Förster, 1869 c g
Teleutaea Förster, 1869 c g
Tetractenion Seyrig, 1932 c g
Tossinola Viktorov, 1958 c g
Tossinolodes Aubert, 1984 c g
Townesion Kasparyan, 1993 c g
Zaglyptomorpha Viereck, 1913 c g
Zygoglypta Momoi, 1965

Data sources: i = ITIS,[9] c = Catalogue of Life,[10] g = GBIF,[11] b =

Townes & Townes (1978)
Brock, J.P. (2017). "The Banchine Wasps (Ichneumonidae: Banchinae) of the British Isles". Handbooks for the Identification of British Insects. Royal Entomological Society. 7 (4): 1–150. ISBN 9781910159019.
Townes (1969, 1971), Townes & Townes (1978), Wahl (1999)
Aubert (1978), Townes & Townes (1978)
Lapointe, Renée; Tanaka, Kohjiro; Barney, Walter E.; Whitfield, James B.; Banks, Jonathan C.; Béliveau, Catherine; Stoltz, Don; Webb, Bruce A.; Cusson, Michel (2007-06-15). "Genomic and Morphological Features of a Banchine Polydnavirus: Comparison with Bracoviruses and Ichnoviruses". Journal of Virology. 81 (12): 6491–6501. doi:10.1128/jvi.02702-06. ISSN 0022-538X. PMC 1900126. PMID 17428854.
Carlson, Robert W. "Superfamily Ichneumonoidea, Discover Life". Retrieved 2018-05-17.
", subfamily Banchinae taxon profile". Retrieved 2018-05-17.
"Classification of Afrotropical Hymenoptera (Wasps, Bees, Ants)". Retrieved 2018-05-17.
"ITIS, Integrated Taxonomic Information System". Retrieved 2018-05-17.
"Catalogue of Life". Retrieved 2018-05-17.

"GBIF". Retrieved 2018-05-17.

Other sources

Aubert, J.F. (1978): Les Ichneumonides ouest-palearctiques et leurs hotes 2. Banchinae et Suppl. aux Pimplinae ["The Western Palearctic ichneumon wasps and their hosts. 2. Banchinae and supplement to the Pimplinae"]. Laboratoire d'Evolution des Etres Organises[verification needed], Paris & EDIFAT-OPIDA, Echauffour. [in French]
Townes, H.K. (1969): Genera of Ichneumonidae, Part 3 (Lycorininae, Banchinae, Scolobatinae, Porizontinae). Memoirs of the American Entomological Institute 13: 1-307.
Townes, H.K. (1971): Genera of Ichneumonidae, Part 4 (Cremastinae, Phrudinae, Tersilochinae, Ophioninae, Mesochorinae, Metopiinae, Anomalinae, Acaenitinae, Microleptinae, Orthopelmatinae, Collyriinae, Orthocentrinae, Diplazontinae). Memoirs of the American Entomological Institute 17: 1-372.
Townes, H.K. & Townes, M. (1978): Ichneumon-flies of America North of Mexico: 7. Subfamily Banchinae, tribes Lissonotini and Banchini. Memoirs of the American Entomological Institute 26: 1-614.
Wahl, David (1999): Classification and Systematics of the Ichneumonidae (Hymenoptera). Version of 1999-JUL-19. Retrieved 2008-JUN-18.

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