Bothynus

Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Endopterygota
Superordo: Coleopterida
Ordo: Coleoptera
Subordo: Polyphaga
Infraordo: Scarabaeiformia
Superfamilia: Scarabaeoidea

Familia: Scarabaeidae
Subfamilia: Dynastinae
Tribus: Pentodontini
Subtribus: Pentodontina
Genus: Bothynus
Subgenera: Oxygrylius

Species (31): B. alvarengai – B. ascanius – B. complanus – B. complexus – B. cribrarius – B. cunctator – B. cyclops – B. cylindricus – B. dasypleurus – B. deiphobus – B. entellus – B. exaratus – B. herbivorus – B. herteli – B. horridus – B. laevipennis – B. lancifer – B. laticifex – B. medon – B. minor – B. niger – B. nyx – B. peninsularis – B. quadridens – B. ruginasus – B. stenelus – B. striatellus – B. thrix – B. tricornis – B. validus – B. villiersi

Bothynus (s. str.) species (28): B. alvarengai – B. ascanius – B. complanus – B. complexus – B. cribrarius – B. cunctator – B. cyclops – B. cylindricus – B. dasypleurus – B. deiphobus – B. entellus – B. exaratus – B. herbivorus – B. herteli – B. horridus – B. laevipennis – B. lancifer – B. laticifex – B. medon – B. minor – B. niger – B. nyx – B. quadridens – B. stenelus – B. striatellus – B. tricornis – B. validus – B. villiersi
Name

Bothynus Hope, 1837: 95

Type species:

References
Primary references

Hope, F.W., 1837: The coleopterist’s manual, containing the Lamellicorn insects of Linneus and Fabricius. Henry G. Bohn, London, pp. i-xiii + 15–121 + [1], pls. 1–3. [original description: p. 95]

Additional references

Hardy, M. 2017. Description of a new species of Bothynus Hope from Argentina and Bolivia (Coleoptera: Scarabaeidae: Dynastinae). Zootaxa 4362(1): 141–145. DOI: 10.11646/zootaxa.4362.1.9 Full article (PDF) Reference page.
Ratcliffe, B.C. 2010: A new Bolivian Bothynus Hope, 1837 (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini), with a key to the species in Bolivia. Coleopterists bulletin, 64: 105–108. DOI: 10.1649/072.064.0201
Ratcliffe, B.C.; Smith, A.B.T. 2007: Case 3369. Bothynus Hope, 1837 (Insecta, Coleoptera, SCARABAEIDAE): proposed conservation of usage by designation of Scarabaeus ascanius Kirby, 1819 as the type species. Bulletin of zoological nomenclature, 64: 29–31. [1]

Links

Schoolmeesters, P. 2017. Scarabs: World Scarabaeidae Database (version Jul 2016). In: Roskov Y., Abucay L., Orrell T., Nicolson D., Bailly N., Kirk P., Bourgoin T., DeWalt R.E., Decock W., De Wever A., Nieukerken E. van, Zarucchi J., Penev L., eds. 2017. Species 2000 & ITIS Catalogue of Life, 30th January 2017. Digital resource at www.catalogueoflife.org/col. Species 2000: Naturalis, Leiden, the Netherlands. ISSN 2405-8858. Reference page. [accessed on Jun 7, 2017]