Bruchinae

Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Endopterygota
Superordo: Coleopterida
Ordo: Coleoptera
Subordo: Polyphaga
Infraordo: Cucujiformia
Cladus: Phytophaga
Superfamilia: Chrysomeloidea

Familia:Chrysomelidae
Subfamilia: Bruchinae
Tribus (6 + 1†): Amblycerini – Bruchini – Eubaptini – Kytorhinini – Pachymerini – Rhaebini – †Myanmaropini
Source(s) of checklist:

Source(s) of checklist:
Checklist based uncritically on Morse (2014), plus Myanmaropini.

Overview of genera (70 + 4†)

Abutiloneus – Acanthobruchidius – Acanthoscelides – Afroredon – Algarobius – Altheus – Amblycerus – Bonaerius – Borowiecus – Bruchidius – Bruchus – Buburra – Butiobruchus – Callosobruchus – Caryedes – Caryedon – Caryoborus – Caryobruchus – Caryopemon – Caryotrypes – Conicobruchus – Cosmobruchus – Ctenocolum – Dahlibruchus – Decellebruchus – Diedobruchus – Eubaptus – Exoctenophorus – Gibbobruchus – Horridobruchus – Kingsolverius – Kytorhinus – Lithraeus – Margaritabruchus – Megabruchidius – Megacerus – Meganeltumius – Meibomeus – Merobruchus – Mimocaryedon – Mimosestes – Neltumius – Neobruchidius – Pachymerus – Palaeobruchidius – Paleoacanthoscelides – Palpibruchus – Parasulcobruchus – Pectinibruchus – Penthobruchus – Protobruchus – Protocaryopemon – Pseudopachymerina – Pygiopachymerus – Pygobruchidius – Rhaebus – Rhipibruchus – Salviabruchus – Scutobruchus – Sennius – Serratobruchidius – Spatulobruchus – Speciomerus – Specularius – Spermophagus – Stator – Stylantheus – Sulcobruchus – Tuberculobruchus – Zabrotes – †Electrocaryedon – †Mesopachymerus – †Myanmarops – †Oligobruchus
Name

Bruchinae Latreille, 1802 [conserved name]

Type genus: Bruchus Linnaeus, 1767

Placed on the Official List of Family-Group Names in Zoology by Opinion 1809. [as "Bruchidae"]
Synonymy

Bruchelae Latreille, 1802: 192
Bruchidae: Stephens, 1831: 206
Bruchinae
Bruchetae Rosenhauer, 1856: 233 [syn.: Yus-Ramos, 2014: 342]
Mylabridae Heyden, 1883
Lariidae Bedel, 1901

Alternative classifications
As family

Family Bruchidae
Subfamily Amblycerinae
Tribe Amblycerini
Tribe Spermophagini
Subfamily Bruchinae
Tribe Acanthoscelidini
Tribe Bruchidiini
Tribe Bruchiini
Tribe Megacerini
Subfamily Eubaptinae
Subfamily Kytorhininae
Subfamily Pachymerinae
Tribe Caryedontini
Tribe Caryopemini
Tribe Pachymerini
Subfamily Rhaebinae

References
Primary references

ICZN. 1995. Opinion 1809. Bruchus Linnaeus, 1767, Ptinus Linnaeus, 1767 and Mylabris Fabricius, 1775 (Insecta: Coleoptera): conserved. Bulletin of Zoological Nomenclature 52(2): 208–210. BHL. Reference page.
Latreille, P.A. 1802. Histoire naturelle, générale et particulière des crustacés et des insectes. Ouvrage faisant suite à l’histoire naturelle générale et particulière, composée par Leclerc de Buffon, et rédigée par C.S. Sonnini, membre de plusieurs sociétés savantes. Familles naturelles des genres. Tome troisième. F. Dufart, Paris, xii + pp. 13–467 + [1 (errata)]. BHL Reference page.
Rosenhauer, W.G. 1856. Die Thiere Andalusiens nach dem Resultate einer Reise zusammengestellt, nebst den Beschreibungen von 249 neuen oder bis jetzt noch unbeschriebenen Gattungen und Arten. Erlangen: T. Blaesing, viii + 429 pp., 3 pls. BHL Reference page.
Stephens, J.F. 1831–1832. Illustrations of British entomology; or, a synopsis of indigenous insects: containing their generic and specific distinctions; with an account of their metamorphoses, times of appearance, localities, food, and economy, as far as practicable. Mandibulata, Vol. IV. London: Baldwin & Cradock, 413 pp., pls. 20–23. [published in parts: pp. 1–366, 1831; 367–413, 1832] BHL Reference page.
Yus-Ramos, R. 2014. Nueva sinonimia de la familia Bruchidae y comentarios sobre el origen de este nombre de familia (Coleoptera). Boletín de la Asociación Española de Entomología 38(3-4): 341–349. Online (with PDF download).

Additional references

Anton, K.-W. 2010. Subfamily Bruchinae Latreille, 1802. Pp. 339–353. In: Löbl, I. & Smetana, A. (eds.) Catalogue of Palaearctic Coleoptera. Volume 6. Chrysomeloidea. Apollo Books, Stenstrup, Denmark. 924 pp. ISBN 978-87-88757-84-2. Google Books. Reference page.
Borowiec, L. 1987. The genera of seed-beetles (Coleoptera, Bruchidae). Polskie Pismo Entomologiczne 57: 3–207. PDF. Reference page.
Ghahari, H. & Borowiec, L. 2017. A checklist of seed-beetles (Coleoptera: Chrysomelidae: Bruchinae) from Iran. Zootaxa 4268(2): 215–237. DOI: 10.11646/zootaxa.4268.2.3 Paywall. ResearchGate Open access. Reference page.
Johnson, C.D. & Romero-Nápoles, J. 2006. Neobruchidius lovie, new genus and new species from Latin America (Coleoptera: Chrysomelidae: Bruchinae). Zootaxa 1123: 57–68. Abstract & excerpt. DOI: 10.11646/zootaxa.1123.1.4 Paywall. PDF. Reference page.
Kingsolver, J.M. 1965. A new fossil bruchid genus and its relationships to modern genera (Coleoptera: Bruchidae: Pachymerinae). The Coleopterists Bulletin 19: 25–30. BHL.
Kingsolver, J.M. 2004. Handbook of the Bruchidae of the United States and Canada. Volume 1. [1]
Kingsolver, J.M. 2004. Handbook of the Bruchidae of the United States and Canada. Volume 2 (illustrations). [2]
Legalov, A.A. 2016. A new genus of seed beetles (Coleoptera, Chrysomelidae, Bruchinae) in Baltic amber. Paleontological Journal 50(1): 73–77. DOI: 10.1134/S0031030116010056 Paywall.
Legalov, A.A., Kirejtshuk A.G. & Anokhin, B.A. 2020. The oldest seed beetle (Coleoptera; Chrysomelidae: Bruchinae) from Upper Cretaceous amber of northern Myanmar with description of new tribe, genus and species. Cretaceous Research 107(104283): 1–5. DOI: 10.1016/j.cretres.2019.104283 Paywall. Reference page.
Morse, G. 2014. 2.7.1. Bruchinae Latreille, 1802. Pp. 189–198 in: Leschen et al., 2014. Full chapter 2.7 (with link to PDF) Reference page.
Nilsson, J.A. & Johnson, C.D. 1993. A taxonomic revision of the palm bruchids (Pachymerini) and a description of the world genera of Pachymerinae. Memoirs of the American Entomological Society No. 41. 104 pp. BHL.
Poinar, G., Jr. 2005. A Cretaceous palm bruchid, Mesopachymerus antiqua, n. gen., n. sp. (Coleptera: Bruchidae: Pachymerini) and biogeographical implications. Proceedings of the Entomological Society of Washington 107(2): 392–397. BHL.
Romero, J. & Johnson, C.D. 2004. Checklist of the Bruchidae (Insecta: Coleoptera) of Mexico. The Coleopterists Bulletin 58(4): 613–635. DOI: 10.1649/694 Paywall. JSTOR Hybrid open access journal. ResearchGate Open access green.
Viana, J.H. & Ribeiro-Costa, C.S. 2013 (Online) 2014 (Print). Bruchines (Coleoptera: Chrysomelidae) associated with Senna neglecta (Vogel) H.S. Irwin and Barneby (Fabaceae: Caesalpinioideae): a new host plant for the subfamily. Journal of Natural History 48(1–2): 57–85. DOI: 10.1080/00222933.2013.791882 Paywall. Reference page.
Yus Ramos, R. 2013. Serratobruchidius, un género nuevo de Bruchidiini para los Bruchidius Schilsky (s.l.) del grupo serraticornis (Coleoptera: Bruchidae). Boletín de la Sociedad Entomológica Aragonesa 52: 17–21. PDF Reference page.
Yus-Ramos, R., Ventura, D., Bensusan, K., Coello-García, P., György, Z. & Stojanova, A. 2014. Alien seed beetles (Coleoptera: Chrysomelidae: Bruchinae) in Europe. Zootaxa 3826(3): 401–448. DOI: 10.11646/zootaxa.3826.3.1 Reference page.
Zampetti, M.F. & Toma, L. 2020. Description of seven new species of seed beetles (Coleoptera Bruchidae) from Mediterranean basin and Africa. Biodiversity Journal 11(2): 553–564. DOI: 10.31396/Biodiv.Jour.2020.11.2.553.564 Open access. [as Bruchidae]

Links

BRUCHBASE by (Geoff Morse)
Bruchidae in SYNOPSIS OF THE DESCRIBED COLEOPTERA OF THE WORLD
Atlas of seed beetles (Bruchidae) of Russia and adjacent countries

Vernacular names
Deutsch: Samenkäfer
English: Bean Weevils / Seed Beetles
français: Bruche
magyar: Zsizsikformák
日本語: マメゾウムシ亜科
polski: Strąkowcowate
русский: Зерновки

The bean weevils or seed beetles are a subfamily (Bruchinae) of beetles, now placed in the family Chrysomelidae, though they have historically been treated as a separate family. They are granivores, and typically infest various kinds of seeds or beans, living most of their lives inside a single seed. The subfamily includes about 1,650 species and are found worldwide.

Bean weevils are generally compact and oval in shape, with small heads somewhat bent under. Sizes range from 1 to 22 mm for some tropical species. Colors are usually black or brown, often with mottled patterns. Although their mandibles may be elongated, they do not have the long snouts characteristic of true weevils.

Adults deposit eggs on seeds, then the larvae chew their way into the seed. When ready to pupate, the larvae typically cut an exit hole, then return to their feeding chamber. Adult weevils have a habit of feigning death and dropping from a plant when disturbed.

Host plants tend to be legumes, but species will also be found in Convolvulaceae, Arecaceae, and Malvaceae, and several species are considered pests.

One characteristic of the beetles which can be seen in the photo is that the elytra are short, not quite reaching the tip of the abdomen.

Several species are native to Great Britain, but there are also records of several introduced species from stored products in warehouses and dwellings, although these species cannot proliferate outside of heated buildings in that climate.
Contents

1 Genera
2 See also
3 Notes
4 References
5 External links

Genera

This list of genera uses the new classification, treating the bean weevils as a subfamily with six extant tribes, eight or nine subtribes, and one extinct tribe. The former names and ranks for the tribes and subtribes under the old classification, treating the been weevils as a family, are given in parentheses (except for Myanmaropini, which was established only for the new classification).[5][6]

Tribe Amblycerini Bridwell, 1932 (= Subfamily Amblycerinae)
Subtribe Amblycerina Bridwell, 1932 (= Tribe Amblycerini)
Amblycerus Thunberg, 1815
Subtribe Spermophagina Borowiec, 1987 (= Tribe Spermophagini)
Spermophagus Schoenherr, 1833
Zabrotes Horn, 1885
Tribe Bruchini Latreille, 1802 (= Subfamily Bruchinae)
Subtribe Acanthoscelidina Bridwell, 1946 (= Tribe Acanthoscelidini)
Abutiloneus Bridwell, 1946
Acanthoscelides Schilsky, 1905
Algarobius Bridwell, 1946
Althaeus Bridwell, 1946
Bonaerius Bridwell, 1952
Caryedes Hummel, 1827
Cosmobruchus Bridwell, 1931
Ctenocolum Kingsolver & Whitehead, 1974
Dahlibruchus Bridwell, 1931
Gibbobruchus Pic, 1913
Lithraeus Bridwell, 1952
Margaritabruchus Romero & Johnson, 2001
Meibomeus Bridwell, 1946
Megasennius Whitehead & Kingsolver, 1975
Merobruchus Bridwell, 1946
Mimosestes Bridwell, 1946
Neltumius Bridwell, 1946
Neobruchidius Johnson & Romero, 2006
Palpibruchus Borowiec, 1987
Pectinibruchus Kingsolver, 1967
Penthobruchus Kingsolver, 1973
Pseudopachymerina Zacher, 1952
Pygiopachymerus Pic, 1911
Rhipibruchus Bridwell, 1932
Scutobruchus Kingsolver, 1968
Sennius Bridwell, 1946
Spatulobruchus Borowiec, 1987
Stator Bridwell, 1946
Stylantheus Bridwell, 1946
Subtribe Bruchidiina Bridwell, 1946 (= Tribe Bruchidiini)[note 1]
Acanthobruchidius Borowiec, 1980
Borowiecus Anton, 1994
Bruchidius Schilsky, 1905
Callosobruchus Pic, 1902
Conicobruchus Decelle, 1951
Decellebruchus Borowiec, 1987
Horridobruchus Borowiec, 1984
Kingsolverius Borowiec, 1987
Megabruchidius Borowiec, 1984
Palaeoacanthoscelides Borowiec, 1985
Parasulcobruchus Anton, 1999
Pygobruchidius Pic, 1951
Salviabruchus Decelle, 1982
Specularius Bridwell, 1938
Sulcobruchus Chûjô, 1937
Tuberculobruchus Decelle, 1951
Subtribe Bruchina Latreille, 1802 (= Tribe Bruchini)
Bruchus Linnaeus, 1767
Subtribe Megacerina Bridwell, 1946 (= Tribe Megacerini)
Megacerus Fåhraeus, 1839
Tribe Eubaptini Bridwell, 1932 (= Subfamily Eubaptinae)
Eubaptus Lacordaire, 1945
Tribe Kytorhinini Bridwell, 1832 (= Subfamily Kytorhininae)
Kytorhinus Fischer von Waldheim, 1809
Tribe Pachymerini Bridwell, 1929 (= Subfamily Pachymerinae)
Subtribe Caryedontina Bridwell, 1929 (= Tribe Caryedontini)
Aforedon Decelle, 1965
Caryedon Schoenherr, 1823
Caryotrypes Decelle, 1968
Exoctenophorus Decelle, 1968
Mimocaryedon Decelle, 1968
Subtribe Caryopemina Bridwell, 1929 (= Tribe Caryomepini)
Caryopemon Jekel, 1855
Diedobruchus Pic, 1913
Protocaryopemon Borowiec, 1987
Subtribe Pachymerina Bridwell, 1929 (= Tribe Pachymerini)
Butiobruchus Prevett, 1966
Caryobruchus Bridwell, 1929
Caryoborus Schoenherr, 1833
Pachymerus Thunberg, 1805
†Mesopachymerus Poinar, 2005 Canadian amber, Campanian
Tribe Rhaebini Chapuis, 1874 (= Subfamily Rhaebinae)
Rhaebus Fischer von Waldheim, 1824
Tribe †Myanmaropini Legalov et al., 2020
†Myanmarops Legalov et al., 2020[2] Burmese amber, Myanmar, Cenomanian