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Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Endopterygota
Superordo: Coleopterida
Ordo: Coleoptera
Subordo: Polyphaga
Infraordo: Cucujiformia
Superfamilia: Coccinelloidea

Familia: Coccinellidae
Subfamilia: Coccinellinae
Tribus [list of tribus after Sasaji (1971a)]: CoccinelliniDiscotominiHalyziini

Tribus [list of tribus after Kovár (1996)]: CoccinelliniTytthaspidiniDiscotominiHalyziiniSinghikalini

Tribus [list of tribus after Ślipiński (2007)]: CocciduliniChilocoriniCoccinelliniDiominiEpilachniniNoviiniScymnilliniSticholotidiniTelsimiini

Tribus [list of tribus after Seago et al. (2011)]: Argentipilosini – Aspidimerini – Cephaloscymnini – Chilocorini – Coccidulini – Coccinellini – Cryptognathini – Diomini – Epilachnini – Hyperaspidini – Limnichopharini – Monocorynini – Noviini – Ortaliini – Platynaspidini – Plotinini – Shirozuellini – Sticholotidini – Telsimiini

Tribus [list of tribus after Che et al. (2021)]: Stethorini – Coccinellini – Chnoodini – Ortaliini – EpilachniniNoviini – Scymnini – Cryptognathini – Cephaloscymnini – Platynaspidini – Aspidimerini – Hyperaspidini – Brachiacanthini – Diomini – Poriini – Pentiliini – Azyini – Argentipilosini – Chilocorini – Plotinini – Sumniini – Sticholotidini – Cranophorini – Coccidulini – Shirozuellini – Scymnillini


Coccinellinae Latreille, 1807
Primary references

Latreille, P.A. 1807. Genera crustaceorum et insectorum secundem ordinem naturalem in familias disposita, iconibus exemplisque plurimus explicata. Tomus tertius. Parisiis et Argentorati: A. Koenig, 258 pp. BHL Reference page.

Additional references

Che, L.H., Zhang, P., Deng, S.H., Escalona, H.E., Wang, X.M., Li, Y., Pang, H., Vandenberg, N., Ślipiński, A., Tomaszewska, W., Liang, D. 2021. New insights into the phylogeny and evolution of lady beetles (Coleoptera: Coccinellidae) by extensive sampling of genes and species. Molecular Phylogenetics and Evolution 156: 107045. DOI: 10.1016/j.ympev.2020.107045 Paywall. ResearchGate [Accepted manuscript] Open access. Reference page.
Kovár, I. 1996: Phylogeny. in: Hodek, I.; Honek, A. (eds), Ecology of Coccinellidae. Kluwer Academic Publishers, Dordrecht: 19–31.
Nedvěd, O. & Kovář, I., 2012. Appendix: List of Genera in Tribes and Subfamilies, pp. 526-531. In: Hodek, I., van Emden, H.F. & Honěk, A. (Eds.) Ecology and Behaviour of the Ladybird Beetles (Coccinellidae), First Edition. Blackwell Publishing Ltd. 561pp. ISBN 9781405184229 DOI: 10.1002/9781118223208 Full text Reference page.
Sasaji, H. 1971a: Fauna Japonica. Coccinellidae (Insecta: Coleoptera). Academic Press of Japan, Keigaku Publishing, Tokyo.
Seago, A.E., Giorgi, J.A., Li, J-H. & Ślipiński, A. 2011. Phylogeny, classification and evolution of ladybird beetles (Coleoptera: Coccinellidae) based on simultaneous analysis of molecular and morphological data. Molecular Phylogenetics and Evolution 60: 137–151. DOI: 10.1016/j.ympev.2011.03.015 Paywall. Full text PDF Reference page.
Ślipiński, S.A. 2007. Australian ladybird beetles (Coleoptera: Coccinellidae). Their biology and classification. Australian Biological Resources Study, Canberra. ISBN 9780642568557. Google books (preview) Reference page.


Helmut Fürsch. Taxonomy of Coccinellids;
Atlas of ladybirds (Coccinellidae) of Russia

Vernacular names
日本語: テントウムシ亜科
中文: 瓢蟲亞科

Coccinellinae is a subfamily of lady beetles in the family Coccinellidae. There are at least 20 genera and 90 described species in Coccinellinae.[1][2][3]

1 Genera
2 Gallery
3 References
4 Further reading
5 External links

Further information: List of Coccinellidae genera § Subfamily Coccinellinae

These 29 genera belong to the subfamily Coccinellinae:

Adalia Mulsant, 1846 i c g b
Aiolocaria (Crotch, 1871)g
Anatis Mulsant, 1846 i c g b (giant lady beetles)
Anisosticta Chevrolat in Dejean, 1837 i c g b
Aphidecta Weise, 1893 i c g b
Calvia Mulsant, 1850 i c g b
Ceratomegilla Crotch, 1873 i c g b
Cheilomenes Chevrolat in Dejean, 1837 i c g
Coccinella Linnaeus, 1758 i c g b
Coelophora Mulsant, 1850 i c g b
Coleomegilla Timberlake, 1920 i c g b
Cycloneda Crotch, 1871 i c g b (spotless lady beetles)
Harmonia Mulsant, 1850 i c g b
Hippodamia Chevrolat in Dejean, 1837 i c g b
Macronaemia Casey, 1899 i c g b
Megalocaria Crotch, 1871 i c g
Micraspis Chevrolat in Dejean, 1837 i c g
Mulsantina Weise, 1906 i c g b
Myzia Mulsant, 1846 i c g b
Naemia Mulsant, 1850 i c g b
Neda Mulsant, 1850 g b
Neoharmonia Crotch, 1871 i c g b
Olla Casey, 1899 i c g b
Paranaemia Casey, 1899 i c g b
Propylea Mulsant, 1846 g b
Psyllobora Chevrolat in Dejean, 1837 i c g b (fungus-eating lady beetles)
Synonycha Chevrolat in Dejean, 1837 i c g
Verania Mulsant, 1850 i c g

Data sources: i = ITIS,[1] c = Catalogue of Life,[4] g = GBIF,[5] b =[2]


"Coccinellinae Report". Integrated Taxonomic Information System. Retrieved 2018-04-09.
"Coccinellinae Subfamily Information". Retrieved 2018-04-09.
"Coccinellinae Overview". Encyclopedia of Life. Retrieved 2018-04-09.
"Catalogue of Life". Retrieved 2018-04-09.

"GBIF". Retrieved 2018-04-09.

Further reading

Arnett, R.H. Jr.; Thomas, M. C.; Skelley, P. E.; Frank, J. H., eds. (2002). American Beetles, Volume II: Polyphaga: Scarabaeoidea through Curculionoidea. CRC Press. ISBN 978-0849309540.
Blatchley, W.S. (1910). An illustrated descriptive catalogue of the Coleoptera, beetles (exclusive of the Rhynchophora) known to occur in Indiana. Nature Pub.
Bouchard, Patrice; Bousquet, Yves; Davies, Anthony E.; Alonso-Zarazaga, Miguel A.; et al. (2011). "Family-group names in Coleoptera (Insecta)". ZooKeys. Pensoft Publishers (88): 1–972. doi:10.3897/zookeys.88.807. ISSN 1313-2989. PMC 3088472. PMID 21594053.
Casey, Thomas L. (1908). "Notes on the Coccinellidae". The Canadian Entomologist. 40 (11): 393–421. doi:10.4039/Ent40393-11.
Crotch, G.R. (1873). Check list of the Coleoptera of America, north of Mexico. Naturalists' Agency. doi:10.5962/bhl.title.38811. ISBN 0665070772.
Evans, Arthur V.; Hogue, James N. (2006). Field Guide to Beetles of California. University of California Press. ISBN 978-0520246577.
Gordon, Robert D. (1985). "The Coccinellidae (Coleoptera) of America North of Mexico". Journal of the New York Entomological Society. 93 (1): 1–912. ISSN 0028-7199.
Gordon, Robert; Vandenberg, Natalia (1991). "Field guide to recently introduced species of Coccinellidae (Coleoptera) in North America". Proceedings of the Entomological Society of Washington.
Klimaszewski, Jan; Watt, J.Charles (1997). "Coleoptera: family-group review and keys to identification". Fauna of New Zealand. 37.
Lawrence, J.F.; Newton Jr., A.F. (1995). Pakaluk, James; Slipinski, Stanislaw Adam (eds.). Families and subfamilies of Coleoptera (with selected genera, notes, references and data on family-group names). Biology, Phylogeny, and Classification of Coleoptera: Papers Celebrating the 80th Birthday of Roy A. Crowson. Vol. 2. Muzeum i Instytut Zoologii PAN. pp. 779–1006. ISBN 83-85192-34-4.
LeConte, J.L. (1861). Classification of the Coleoptera of North America. Smithsonian Miscellaneous Collections. Vol. 3. Smithsonian Institution. doi:10.5962/bhl.title.38459. ISBN 0665100558.
Leng, Charles W. (1920). Catalogue of the Coleoptera of America, North of Mexico. John D. Sherman, Jr. doi:10.5962/bhl.title.8777.
Poole, Robert W.; Gentili, Patricia, eds. (1996). "A Check List of the Insects of North America: Coleoptera". Nomina Insecta Nearctica: A Check List of the Insects of North America. Entomological Information Services. 1: Coleoptera, Strepsiptera: 41–820. ISBN 9781889002019.
Vandenberg, Natalia J. (2002). Arnett, Ross H. Jr.; Thomas, M.C.; Skelley, P.E.; Frank, J.H. (eds.). Family 93; Coccinellidae Latreille 1807. American Beetles. Vol. 2: Polyphaga: Scarabaeoidea through Curculionoidea. CRC Press. pp. 371–389. ISBN 0-8493-0954-9.
White, Richard E. (1998) [1983]. A Field Guide to the Beetles of North America (Peterson Field Guides). Houghton Mifflin Harcourt. ISBN 0395910897.

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