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Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Endopterygota
Superordo: Coleopterida
Ordo: Coleoptera
Subordo: Polyphaga
Infraordo: Cucujiformia
Superfamilia: Coccinelloidea

Familia: Endomychidae
Subfamiliae: Cyclotominae – DanascelinaeEndomychinaeEpipocinaeLeiestinaeLycoperdininaeMerophysiinaePleganophorinae – Xenomycetinae

Overview of genera

Eumorphus – Cholovocera – ...

Endomychidae Leach, 1815



Leach, W.E. 1815. Entomology [pp. 57–172]. In: Brewster, D. (Ed). Brewster’s Edinburgh Encyclopedia. Volume IX [part I]. W. Blackwood, J. Waugh, etc., Edinburgh, 764 pp. BHL Reference page.
I.Löbl & A.Smetana (eds). 2007 Catalogue of Palearctic Coleoptera. Vol. 4: Elateroidea, Derodontoidea, Bostrichoidea, Lymexyloidea, Cleroidea and Cucujoidea. Apollo Books, Stenstrup, Denmark ISBN 87-88757-67-6, p. 557
Majka, C.G. 2009: Erratum: New Brunswick Endomychidae (Coleoptera). Zootaxa, 2160: 68. PDF
Robertson, J.A., Ślipiński, A., Moulton, M., Shockley, F.W., Giorgi, J.A., Lord, N.P., McKenna, D.D., Tomaszewska, W., Forrester, J., Miller, K.B., Whiting, M.F. & McHugh, J.V. 2015. Phylogeny and classification of Cucujoidea and the recognition of a new superfamily Coccinelloidea (Coleoptera: Cucujiformia). Systematic Entomology 40(4): 745–778. DOI: 10.1111/syen.12138 Paywall. ResearchGate Open access. Reference page.
Shockley, F.W. 2009: Family Endomychidae. Pp. 109-114 in: Gerlach, J. (ed.) The Coleoptera of the Seychelles Islands. Pensoft Publishers, Sofia-Moscow. ISBN 9789546424983
Shockley, F.W.; Tomaszewska, K.W.; McHugh, J.V. 2009: An annotated checklist of the handsome fungus beetles of the world (Coleoptera: Cucujoidea: Endomychidae). Zootaxa, 1999: 1–113. Abstract & excerpt
Shockley, F.W.; Tomaszewska, K.W.; McHugh, J.V. 2009: Review of the natural history of the handsome fungus beetles (Coleoptera: Cucujoidea: Endomychidae). Insecta Mundi, 0072: 1–24. PDF
Shockley, F.W.; Vandenberg, N.J. 2009: Catalogue of the primary types of Cerylonidae, Endomychidae and Latridiidae (Coleoptera: Cucujoidea) deposited in the National Museum of Natural History, with additional notes and clarification of the status of several types. Zootaxa, 2229: 1–64. Abstract & excerpt
Tomaszewska, K.W. 2000: Morphology, phylogeny and classification of adult Endomychidae (Coleoptera: Cucujoidea). Annales Zoologici, 50: 449–558. Abstract PDF
Tomaszewska, W. 2010: 10.32. Endomychidae Leach, 1815. Pp. 442-454 in: Leschen, R.A.B.; Beutel, R.G.; Lawrence, J.F. (volume eds.) Coleoptera, beetles. Volume 2: Morphology and systematics (Elateroidea, Bostrichiformia, Cucujiformia partim). In: Kristensen, N.P. & Beutel, R.G. (eds.) Handbook of zoology. A natural history of the phyla of the animal kingdom. Volume IV. Arthropoda: Insecta. Part 38. Berlin, New York: Walter de Gruyter. ISBN 3110190753 ISBN 9783110190755
Yoshitomi, H. & Sogoh, K. 2019. Eumorphus marginatus group in Sulawesi, Indonesia (Coleoptera, Endomychidae). Zookeys, 820: 139–149. DOI: 10.3897/zookeys.820.31527 Reference page.


Australian Faunal Directory (stub)
Shockley, Floyd W. 2009. Endomychidae. Handsome fungus beetles. Version 31 March 2009 in The Tree of Life Web Project

Vernacular names
English: handsome fungus beetles

Endomychidae, or handsome fungus beetles, is a family of beetles with representatives found in all biogeographic realms. There are around 120 genera and 1300 species. The family was established based on the type genus Endomychus, a genus erected in 1795 by Panzer which was applied to a species that Linnaeus called Chrysomela coccinea. As the common name suggests, Endomychidae feed on fungi. Crowson, in his influential treatment of the beetles, placed the family within the Cucujoidea. They have a tarsal formal of 4-4-4 or 3-3-3 and the wings lack a closed radial cell. The second antennal segment has a sensory appendage that is as long as the third antennal segment. The family has also been grouped with the Coccinellidae in a group called the Trimera for having pseudotrimerous tarsi.[1] A 2015 molecular phylogeny study found that the Cucujoidea were found to be non-monophyletic and the Endomychidae was refined with the removal of the Anamorphinae from within the family and elevated to the status of a full family, Anamorphidae. Mycetaeinae and Eupsilobiinae were also found not to belong within the clades of the core Endomychidae, and likewise reclassified into the families Mycetaeidae and Eupsilobiidae.[2]

The subfamilies that are included:

Endomychinae (including Stenotarsinae)


These 70 genera belong to the family Endomychidae as defined in the past (due to revisions, this may not be entirely accurate):

Acinaces Gerstaecker, 1858 g
Aclemmysa Reitter, 1904 g
Agaricophilus Motschulsky, 1838 g
Amphisternus Germar, 1843 g
Amphistethus Strohecker, 1964 g
Ancylopus Costa, 1854 g
Anidrytus Gerstaecker, 1858 i c g
Aphorista Gorham, 1873 i c g b
Archipines Strohecker, 1953 i c g
Asymbius Gorham, 1896 g
Atopomychus Tomaszewska & Szawaryn, 2013 g
Atrichonota Arrow, 1925 g
Avencymon Strohecker, 1971 g
Beccariola Arrow, 1943 g
Bolbomorphus Gorham, 1888 g
Brachytrycherus Arrow, 1920 g
Bystodes Strohecker, 1953 g
Cholovocera Motschulsky, 1838 g
Chondria Gorham, 1888 g
Corynomalus Chevrolat in Dejean, 1836 i c g
Cyclotoma Mulsant, 1851 i g
Danae Reiche, 1847 i c g b
Danascelis Tomaszewska, 1999 i c g
Dapsa Latreille, 1829 g
Dexialia Sasaji, 1970 g
Dialexia Gorham, 1891 g
Discolomopsis Shockley, 2006 g
Displotera Reitter, 1877 g
Ectomychus Gorham, 1888 g
Eidorus g
Endomychus Panzer, 1795 i c g b
Ephebus Chevrolat in Dejean, 1836 i c g
Epipocus Germar, 1843 i c g b
Epopterus Chevrolat in Dejean, 1836 i c g
Eucteanus Gerstaecker, 1857 g
Eucymon g
Eumorphus Weber, 1801 g
Geoendomychus Lea, 1922 g
Glesirhanis Shockley & Alekseev, 2014 g
Hadromychus Bousquet & Leschen, 2002 i c g b
Holoparamecus Curtis, 1833 i c g b
Hylaia Guerin, 1857 g
Hylaperdina Tomaszewska, 2012 g
Idiophyes Blackburn, 1896 g
Indalmus Gerstaecker, 1858 g
Leiestes Chevrolat, 1836 g
Lycoperdina Latreille, 1807 i c g b
Meilichius Gerstaecker, 1857 g
Merophysia Lucas, 1852 g
Microxenus Wallaston, 1861 g
Mycetina Mulsant, 1846 i c g b
Mychothenus Strohecker, 1953 g
Natalinus Tomaszewska, 2011 g
Ohtaius Chûjô, 1938 g
Palaeoestes Kirejtshuk & Nel, 2009 g
Paniegena Heller, 1916 g
Parasymbius Arrow, 1920 g
Phymaphora Newman, 1838 i c g b
Phymaphoroides Motschoulsky, 1857 g
Pleganophorus Hampe, 1855 g
Polymus Mulsant, 1846 g
Pseudindalmus Arrow, 1920 g
Rhanidea Strohecker, 1953 i c g b
Saula Gerstaecker, 1858 i c g
Sinocymbachus Strohecker & Chujo, 1970 g
Stenotarsus Perty, 1832 i c g b
Stethorhanis Blaisdell, 1931 i c g
Trochoideus Westwood, 1833 i c g b
Trycherus Gerstaecker, 1857 g
Xenomycetes Horn, 1880 i c g b

Data sources: i = ITIS,[3] c = Catalogue of Life,[4] g = GBIF,[5] b =[6]
Fossil genera

Burmalestes Tomaszewska and Ślipiński, 2018[7] Burmese amber, Myanmar, Cenomanian
Cretolestes Tomaszewska, Ślipiński and Ren, 2018[7] Burmese amber, Myanmar, Cenomanian
Cretaparamecus Tomaszewska, Ślipiński, Bai and Zhang, 2018[7] Burmese amber, Myanmar, Cenomanian
Palaeomycetes Tomaszewska, Ślipiński and Ren, 2018[7] Burmese amber, Myanmar, Cenomanian
Zemyna Tomaszewska, 2018[7] (replaced name Laima Alekseev and Tomaszewska, 2018): Baltic amber, Eocene


Tomaszewska, K.Wioletta (2000). "Morphology, phylogeny and classification of adult endomychidae (Coleoptera: Cucujoidea)" (PDF). Annales Zoologici. 50 (4): 449–558.
Robertson, James A; Ślipiński, Adam; Moulton, Matthew; Shockley, Floyd W; Giorgi, Adriano; Lord, Nathan P; McKenna, Duane D; Tomaszewska, Wioletta; Forrester, Juanita; Miller, Kelly B; Whiting, Michael F; McHugh, Joseph V (2015). "Phylogeny and classification of Cucujoidea and the recognition of a new superfamily Coccinelloidea (Coleoptera: Cucujiformia)" (PDF). Systematic Entomology. 40 (4): 745. doi:10.1111/syen.12138.
"Endomychidae Report". Integrated Taxonomic Information System. Retrieved 2018-05-07.
"Browse Endomychidae". Catalogue of Life. Retrieved 2018-05-07.
"Endomychidae". GBIF. Retrieved 2018-05-07.
"Endomychidae Family Information". Retrieved 2018-05-07.
Tomaszewska, Wioletta; Ślipiński, Adam; Bai, Ming; Zhang, Weiwei; Ren, Dong (November 2018). "The oldest representatives of Endomychidae (Coleoptera: Coccinelloidea) from the Upper Cretaceous Burmese amber". Cretaceous Research. 91: 287–298. doi:10.1016/j.cretres.2018.07.001.

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