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Superregnum: Eukaryota
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Endopterygota
Superordo: Panorpida
Cladus: Amphiesmenoptera
Ordo: Lepidoptera
Subordo: Glossata
Cladus: Coelolepida
Cladus: Myoglossata
Cladus: Neolepidoptera
Infraordo: Heteroneura
Cladus: Eulepidoptera
Cladus: Ditrysia
Cladus: Apoditrysia
Cladus: Obtectomera
Superfamilia: Papilionoidea

Familia: Hesperiidae
Subfamilia: Hesperiinae
Tribus: Hesperiini
Subtribus: Hesperiina
Genus: Euphyes
Species: E. ampa – E. antra – E. arpa – E. bayensis – E. berryi – E. bimacula – E. canda – E. chamuli – E. cherra – E. conspicua – E. cornelius – E. dion – E. dukesi – E. eberti – E. fumata – E. leptosema – E. peneia – E. pilatka – E. singularis – E. subferrugineus – E. vestris

Euphyes Scudder, 1872.

Type species: Hesperia metacomet Harris, 1862.

Euphyes Scudder, 1872: 80; Evans, 1955: 362; Lamas, 2004: 67.
Euphiges Keferstein, 1874. Mis-spelling; per Lamas, 2004: 67.
Euphyse Anon. 1971. Mis-spelling; per Lamas, 2004: 67.
Eupheys Ramos, 1982. Mis-spelling; per Lamas, 2004: 67.
Euphes Raguso & Lorente, 1997. Mis-spelling; per Lamas, 2004: 67.

Primary references

Scudder, S.H. 1872. A systematic Revision of some of the American Butterflies; with brief notes on those known to occur in Essex County, Mass. 4th Ann. Rep. Peabody Academy of Sciences (1871): 24–83. Online. Reference page.

Additional references

Evans, W.H. 1951–55. A Catalogue of the American Hesperiidae. Pt.1 (1951): x+92pp, pls. 1–9; Pt.2 (1952): v.+178pp, pls. 10–25; Pt.3 (1953): v+246pp, pls. 26–53; Pt.4 (1955): v+499pp, pls. 54–88. Reference page.
Freeman, H.A. 1969. Records, new species, and a new genus of Hesperiidae from Mexico. Journal of the Lepidopterists' Society 23(suppl. 2): 1–64. Reference page.
Freeman, H.A. 1975. A new species of Euphyes Scudder from Texas (Hesperiidae). Journal of the Lepidopterists' Society 29(4): 227–229. PDF. Reference page.
Gatrelle, R.R. 1999. Three new Hesperioidae (Hesperiinae) from South Carolina: new subspecies of Euphyes bimacula, Poanes aaroni, and Hesperia attalus. The Taxonomic Report 1 (10): 1–13. Full article (PDF). Reference page.
Lamas, G. 2004. (ed.) Checklist: Part 4A. Hesperioidea - Papilionoidea. In Heppner, J.B. (ed.) Atlas of Neotropical Lepidoptera. Vol. 5A. Association for Tropical Lepidoptera/Scientific Publishers, Gainesville. 439p. Reference page.
Li, W., Cong. Q., Shen, J.H., Zhang, J., Hallwachs, W., Janzen, D.H. & Grishin, N.V. 2019. Genomes of skipper butterflies reveal extensive convergence of wing patterns. PNAS 116(13): 6232–6237. DOI: 10.1073/pnas.1821304116. Text. SI Appendix. Reference page.
Miller, L.D., Harvey, D.J. & Miller, J.Y. 1985. Notes on the genus Euphyes, with description of a new subspecies (Lepidoptera: Hesperiidae). Florida Entomologist 68(2): 323–335. PDF. Reference page.
Shuey, J.A. 1988 (1989). The Morpho-Species Concept of Euphyes dion with the Description of a New Species (Hesperiidae). Journal of Research on the Lepidoptera 27(3-4): 160–172. PDF. Reference page.
Shuey, J.A. 1993. Phylogeny and biogeography of Euphyes Scudder (Hesperiidae). Journal of the Lepidopterists' Society 47(4): 261–278. PDF. Reference page.
Steinhauser, S.R. & Warren, A.D. 2001. A new species of Euphyes Scudder from Western Mexico (Hesperiidae: Hesperiinae). Journal of the Lepidopterists' Society 55(4): 158–161. Reference page.


Butterflies of America.

The Euphyes is genus of North American butterfly of the family Hesperiidae (skippers), subfamily Hesperiinae (grass skippers).

Listed alphabetically within species groups.[1][2]

Group peneia:
Euphyes peneia (Godman, [1900]) – guardpost skipper
Euphyes eberti Mielke, 1972 – Ebert's skipper
Euphyes leptosema Mabille, 1891 – leptosema skipper
Euphyes fumata Mielke, 1972 – fumata skipper
Euphyes singularis (Herrich-Schäffer, 1865) – singularis skipper [Butler's branded skipper (E. s. insolata) in Jamaica
Euphyes cornelius (Latreille, [1824]) – Cornelius skipper
Group subferruginea:
Euphyes subferrugineus (Hayward, 1934) – subferrugineus skipper
Euphyes antra Evans, 1955 – Antra skipper
Euphyes cherra Evans, 1955 – Cherra skipper
Group dion:
Euphyes bayensis Shuey, 1989 – bay skipper
Euphyes berryi (E. Bell, 1941) – Berry's skipper
Euphyes conspicua (W. H. Edwards, 1863) – black dash
E. c. buchholzi (P. Ehrlich and Gillham, 1951)[3]
E. c. conspicua (W. H. Edwards, 1863)[3]
E. c. orono (Scudder, 1872)[3]
Euphyes dion (W. H. Edwards, 1879) – Dion skipper
Euphyes dukesi (Lindsey, 1923) – Dukes' skipper
E. d. calhouni Shuey, 1996[4]
E. d. dukesi (Lindsey, 1923)[4]
Euphyes pilatka (W. H. Edwards, 1867) – Palatka skipper or saw-grass skipper
E. p. klotsi L. Miller, Harvey and J. Miller, 1985[5]
E. p. pilatka (W. H. Edwards, 1867)[5]
Group vestris:
Euphyes vestris (Boisduval, 1852) – dun skipper, sedge witch, or dun sedge skipper
E. v. harbisoni J. Brown and Mcguire, 1983[6]
E. v. kiowah (Reakirt, 1866)[6]
E. v. metacomet (T. Harris, 1862)[6]
E. v. vestris (Boisduval, 1852)[6]
Euphyes chamuli Freeman, 1969 – Chamul skipper
Euphyes bimacula (Grote & Robinson, 1867) – two-spotted skipper
E. b. arbotsti Gatrelle, 1999[7]
E. b. bimacula (Grote & Robinson, 1867)[7]
E. b. arbotsti (Dodge, 1872)[7]
Euphyes arpa (Boisduval & Le Conte, [1837])[8] – palmetto skipper
Incertae sedis:
Euphyes ampa Evans, 1955 – ampa skipper
Euphyes canda (Steinhauser & Warren, [2002]) – Candelaria skipper


"Archived copy". Archived from the original on 2010-06-23. Retrieved 2010-06-14. Interactive Listing of Mexican Butterflies 11-VII-08
"Euphyes conspicua". Integrated Taxonomic Information System. Retrieved 1 January 2014.
"Euphyes dukesi". Integrated Taxonomic Information System. Retrieved 1 January 2014.
"Euphyes pilatka". Integrated Taxonomic Information System. Retrieved 1 January 2014.
"Euphyes vestris". Integrated Taxonomic Information System. Retrieved 1 January 2014.
"Euphyes bimacula". Integrated Taxonomic Information System. Retrieved 1 January 2014.
"Euphyes arpa". Integrated Taxonomic Information System. Retrieved 1 January 2014.

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