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Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Endopterygota
Superordo: Panorpida
Cladus: Amphiesmenoptera
Ordo: Lepidoptera
Subordo: Glossata
Cladus: Coelolepida
Cladus: Myoglossata
Cladus: Neolepidoptera
Infraordo: Heteroneura
Cladus: Eulepidoptera
Cladus: Ditrysia
Cladus: Apoditrysia
Cladus: Obtectomera
Superfamilia: Papilionoidea

Subfamilia: Satyrinae
Tribus: Haeterini
Genera: Cithaerias - Dulcedo - Haetera - Pierella - Pseudohaetera

Haeterini Herrich-Schäffer, 1864

The Haeterini are one of the smaller tribes of the Satyrinae in the Nymphalidae (brush-footed butterfly) family. The tribe occurs exclusively in tropical rain forests in the Neotropical realm.

It contains 29 described species and 39 subspecies[a] classified in the following five genera:[b]

Cithaerias Hübner, [1819]
Dulcedo d'Almeida, 1951
Haetera Fabricius, 1807
Pierella Westwood, [1851][c]
Pseudohaetera Brown, 1942


But these estimates remain contentious, and at least 5 more species can be recognized based on the multispecies coalescent model as suggested by Matos-Maraví et al. (2019)
According to TOLweb, Lepidoptera and Some Other Life Forms (Savela), Wikispecies, and Archived 8 December 2010 at the Wayback Machine which all seem to be based on Lamas, G. (2004).

Markku Savela's Pierella page says this is invalid and the correct citation is Herrich-Schäffer, 1865, however no source is given for this conclusion and the LepIndex[1] (among other sources) consider Westwood, [1851] to be correct.


Beccaloni, G.; Scoble, M.; Kitching, I.; Simonsen, T.; Robinson, G.; Pitkin, B.; Hine, A.; Lyal, C., eds. (2003). "Pierella". The Global Lepidoptera Names Index. Natural History Museum.

Lamas, G. (ed), 2004. Atlas of Neotropical Lepidoptera. Checklist: Part 4A Hesperioidea - Papiionoidea. Gainesville: Scientific Publishers/Association of Tropical Lepidoptera.
Matos-Maraví, P., Wahlberg, N., Antonelli, A. & Penz, C.M., 2019. Species limits in butterflies (Lepidoptera: Nymphalidae): reconciling classical taxonomy with the multispecies coalescent. Systematic Entomology, in press. DOI:
Wahlberg, N. & Andrew V.Z.B., 2009. Haeterini Herrich-Schaeffer 1864. Version 2 June 2009 (under construction). Archived 6 June 2011 at the Wayback Machine in The Tree of Life Web Project.

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