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Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Endopterygota
Superordo: Panorpida
Cladus: Amphiesmenoptera
Ordo: Lepidoptera
Subordo: Glossata
Cladus: Coelolepida
Cladus: Myoglossata
Cladus: Neolepidoptera
Infraordo: Heteroneura
Cladus: Eulepidoptera
Cladus: Ditrysia
Cladus: Apoditrysia
Cladus: Obtectomera
Superfamilia: Papilionoidea

Familia: Hesperiidae
Subfamilia: Hesperiinae
Tribus: Hesperiini
Subtribus: Hesperiina
Genus: Hesperia
Species: H. assiniboia – H. attalus – H. colorado – H. comma – H. florinda – H. hiraca – H. hyrmina – H. juba – H. lindseyi – H. meskei – H. metea – H. neglecta – H. nabokovi – H. sassacus – H. uncas – H. viridis – H. woodgatei

Hesperia Fabricius, 1793.

Type species: Papilio comma Linnaeus, 1758.

per Lamas, 2004.

Hesperia Fabricius, 1793: 258.
Pamphila Fabricius, 1807: 287. Type species: Papilio comma Linnaeus, 1758, by subsequent designation by Westwood, 1839: 360.
Diorthosus Rafinesque, 1815: 128.
Steropes Rafinesque, 1815: 128; unavailable ICZN Art.12.2.5.
Phidias Rafinesque, 1815: 128; un-necessary replacement name.
Symmachia Sodovskii, 1837ː ; pre-occ Hübner, [1819]. Invalid replacement name.
Ocytes Scudder, 1872ː 76.
Anthomaster Scudder, 1872ː 78.
Herpena Edwards, 1872ː . Mis-spelling.
Anthomastes Scudder, 1881ː . Mis-spelling.
Phamphila Mabille, 1891ː . Mis-spelling.
Pmaphila Maynard, 1891ː . Mis-spelling.
Pamphala Maynard, 1891ː . Mis-spelling.
Pamphilia Moffat, 1894ː . Mis-spelling.
Hesperis Brunner von Wattenwyl, 1897. Mis-spelling.
Pamhpila Maynard, 1901ː . Mis-spelling.
Pamhila Maynard, 1901ː . Mis-spelling.
Urbicola Tutt, 1905ː
Pamphilus Ihering, 1908ː
Herperia Woodworth, 1913ː . Mis-spelling.
Pamphilv Rocha, 1936ː . Mis-spelling.
Hesperria Justino, 1949ː . Mis-spelling.
Hespina dos Passos, 1964ː . Mis-spelling.
Uribicola Bridges, 1988ː . Mis-spelling.


Bell, E.L. & Comstock, W.P. 1948. A new genus and some new species and subspecies of American Hesperiidae (Lepidoptera, Rhopalocera). American Museum Novitates 1379: 1–23. PDF. Reference page.
Cong, Q., Zhang, J., Shen, J.H. & Grishin, N.V. 2019. Fifty new genera of Hesperiidae (Lepidoptera). Insecta Mundi 0731: 1–56. web. Reference page.
Fabricius, J.C. 1807. In Illiger, J. K. W. Magazin für Insektenkunde 6: 279–289. BHL. Reference page.
Gatrelle, R.R. 1999. Three new Hesperioidae (Hesperiinae) from South Carolina: new subspecies of Euphyes bimacula, Poanes aaroni, and Hesperia attalus. The Taxonomic Report 1 (10): 1–13. Full article (PDF). Reference page.
Korb, S.K., 2013. Clarification of data on the Far Eastern representatives of genus Hesperia ̺Fabricius, 1793 (Lepidoptera: Hesperiidae). Eversmannia, 33: 3–4.
Find all Wikispecies pages which cite this reference.</noinclude>
Linnaeus, C. 1758. Systema Naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Editio Decima, Reformata. Tomus I. Holmiæ (Stockholm): impensis direct. Laurentii Salvii. 824 pp. DOI: 10.5962/bhl.title.542 BHL Reference page.
Plötz, C. 1882–83. Die Hesperiinen-Gattung Hesperia Aut. und ihre Arten. Entomologische Zeitung 1882: 43: 314–344, 436-456; 1883: 44: 26–64, 44: 195–233. Reference page.
Rafinesque, C.S. 1815. Analyse de la nature, ou tableau de l'univers et des corps organisés. Palerme: L'Imprimerie de Jean Barravecchia. 224 pp. BHL Reference page.
Scudder, S.H. 1874. The Species of the Lepidopterous Genus: Pamphila. Memoirs read before the Boston Society of Natural History 2: 341–353, pls. 10,11. BHL Reference page.
P.M. Severns & Severns, D.G., 2005. Hesperia lindseyi mccorklei (Hesperiidae): a new subspecies of skipper from Southwestern Oregon, USA. Journal of the Lepidopterists' Society 59(3): 166–169. PDF. Reference page.
Skinner, H., 1890. Notes on Butterflies found at Cape May, N.J. with description of a new species of Pamphila. Entomological News 1(1): 6-9. Reference page.
Skinner, H., 1890. A New Pamphila. Entomologica Americana 6(7): 138. Reference page.
Skinner, H., 1891. A New Pamphila. Entomological News 2(9): 175. Reference page.
W. Sodovskii, 1837. Etymologische Untersuchungen über die Gattungs-namen der Schmetterlinge, von Dr. W. Sodoffsky in Riga. Bulletin de la Société impériale des naturalistes de Moscou 10(6-8)ː 76-97. Reference page.
Warren, Gott and McGuire, 2021. A review of Hesperia uncas W. H. Edwards, 1863 in Mexico, with the description of a new subspecies (Lepidoptera: Hesperiidae: Hesperiinae). PDF Reference page.
Westwood, J.O. 1839. An introduction to the modern classification of insects. Vol. 1. Longman, Orme, Brown, Green, and Longmans, London. 462 pp. BHL Reference page.

Hesperia, the branded skippers, is a Holarctic genus in the skippers (Hesperiidae) butterfly family. Most species are endemic to North America, Hesperia comma is widespread throughout the region. H. florinda is endemic to temperate eastern Asia. H. nabokovi is endemic to Hispaniola.

Presumably Johan Christian Fabricius named the genus for Hesperia, one of the Hesperides.

Hesperia species are:

Hesperia assiniboia (Lyman, 1892) – Plains skipper
Hesperia attalus (W. H. Edwards, 1871) – dotted skipper – south United States
Hesperia attalus attalus (W. H. Edwards, 1871)
Hesperia attalus nigrescens (Gatrelle, 1999) – dark dotted skipper
Hesperia attalus slossonae (Skinner, 1871) – Slosson's dotted skipper
Hesperia caucasica Riabov, 1926 Caucasus
Hesperia colorado (Scudder, 1874) – western branded skipper
Hesperia colorado colorado (Scudder, 1874)
Hesperia colorado mattoonorum (McGuire, 1998)
Hesperia colorado ochracea – ochre branded skipper
Hesperia colorado oregonia – Oregon branded skipper
Hesperia colorado oroplata
Hesperia colorado susanae – Susan's branded skipper
Hesperia columbia (Scudder, 1872) – Columbian skipper – California to Southwest Oregon
Hesperia comma (Linnaeus, 1758) – silver-spotted skipper or common branded skipper
Hesperia dacotae (Skinner, 1911) – Dakota skipper
Hesperia florinda (Butler, 1878)
Hesperia florinda florinda (Butler, 1878) Transbaikalia, Korea, Amur, Ussuri, Japan
Hesperia florinda rozhkovi Kurentzov, 1970
Hesperia juba (Scudder, 1872) – Juba skipper
Hesperia leonardus (Harris, 1862) – Leonard's skipper
Hesperia leonardus leonardus (Harris, 1862) – (= stallingsi (HA Freeman, 1943) – morph)
Hesperia leonardus montana (Skinner, 1911) – mountain skipper
Hesperia leonardus pawnee (Dodge, 1874) – Pawnee skipper (= ogallala (Leussler, 1921))
Hesperia lindseyi (Holland, 1930) – Lindsey's skipper – Southwest Oregon, California, Southwest Arizona
Hesperia lindseyi lindseyi (Holland, 1930)
Hesperia lindseyi eldorado (J. Emmel, T. Emmel & Matoon, 1998)
Hesperia lindseyi macneilli (J. Emmel, T. Emmel & Matoon, 1998)
Hesperia lindseyi septentrionalis (J. Emmel, T. Emmel & Matoon, 1998)
Hesperia meskei W. H. Edwards, 1877 – Meke's skipper, Dixie skipper – Texas to Florida
Hesperia meskei meskei W. H. Edwards, 1877
Hesperia meskei pinocayo (Gatrelle and Minno) – Rockland skipper
Hesperia meskei straton (W. H. Edwards, 1881) – Easter Meske's skipper
Hesperia metea Scudder, 1863 – cobweb skipper
Hesperia metea metea (Scudder, 1864)
Hesperia metea intermedia (Gatrelle) – southern cobweb skipper
Hesperia metea licinus (Edwards, 1871) – Licinus cobweb skipper
Hesperia miriamae MacNeill, 1959 – Sierra skipper – Inyo County, California
Hesperia miriamae miriamae MacNeill, 1959
Hesperia miriamae longaevicola (McGuire, 1998)
Hesperia nabokovi (Bell & Comstock, 1948) Hispaniola
Hesperia nevada (Scudder, 1874) – Nevada skipper
Hesperia nevada nevada (Scudder, 1874)
Hesperia nevada sierra Austin, J. Emmel, T. Emmel & Matoon, 1998 – western Nevada skipper
Hesperia ottoe W. H. Edwards, 1866 – Ottoe skipper
Hesperia pahaska Luessler, 1938 – Pahaska skipper
Hesperia pahaska pahaska Luessler, 1938
Hesperia pahaska martini (MacNeill, 1964) – Martin's skipper
Hesperia pahaska williamsi (Lindsey, 1938) – William's skipper
Hesperia pawnee Dodge, 1874 Colorado
Hesperia sassacus Harris, 1862 – Indian skipper
Hesperia sassacus sassacus Harris, 1862
Hesperia sassacus manitoboides (J. Fletcher, 1888) – northwestern Indian skipper
Hesperia sassacus nantahala (Gatrelle & Grkovich) – Carolina Indian skipper
Hesperia uncas W. H. Edwards, 1863 – Uncas skipper
Hesperia uncas uncas W. H. Edwards, 1863 (= ridingsii (Reakirt, 1866))
Hesperia uncas fulvapella (Austin & McGuire, 1998)
Hesperia uncas gilberti MacNeill, 1964
Hesperia uncas giulianii (McGuire, 1998)
Hesperia uncas grandiosa (Austin & McGuire, 1998)
Hesperia uncas lasus (W. H. Edwards, 1881)
Hesperia uncas macswaini MacNeill, 1964 – MacSwain's skipper
Hesperia uncas reeseorum (Austin & McGuire, 1998) – Reeses' Uncas skipper
Hesperia uncas terraclivosa (Austin & McGuire, 1998)
Hesperia viridis (W. H. Edwards, 1883) – green skipper – Mexico, New Mexico
Hesperia woodgatei (R. C. Williams, 1914) – Apache skipper – Arizona, New Mexico, South Texas, North Mexico


Pamphila was a genus of butterflies in the family Hesperiidae. It is now recognised as a junior objective synonym of Hesperia.[1] A list of species formerly identified as Pamphila.
References and external links

Savela, Markku. "Hesperia Fabricius, 1793". Lepidoptera and Some Other Life Forms. Retrieved October 15, 2017.
Images representing Hesperia at Consortium for the Barcode of Life
Systematic list of the butterflies of Norway
Juba Skipper page
Jeff's Butterfly Page
Checklist of Butterflies of Tulare County
University of Colorado Museum
Gatrelle R. A subspecific assessment of the genus Hesperi (Hesperiinae) in eastern north America (Part I: The south)
Butterflies and Skippers of North America
Standardized Common North American Butterfly names
TC-ISBN Main List
Hesperia from Markku Savela's Lepidoptera site.
Lyman Entomological Museum and Research Laboratory


"Pamphila". Butterflies and Moths of the World. Natural History Museum. Retrieved 14 October 2017.

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