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Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Endopterygota
Superordo: Panorpida
Cladus: Amphiesmenoptera
Ordo: Lepidoptera
Subordo: Glossata
Cladus: Coelolepida
Cladus: Myoglossata
Cladus: Neolepidoptera
Infraordo: Heteroneura
Cladus: Eulepidoptera
Cladus: Ditrysia
Cladus: Apoditrysia
Cladus: Obtectomera
Superfamilia: Papilionoidea

Familia: Lycaenidae
Subfamilia: Miletinae
Tribus (5): Lachnocnemini - Liphyrini - Miletini - Spalgini - Tarakini


Miletinae Corbet, 1939: 63.
Type Genus: Miletus Hübner, 1819: 71.

Gerydinae Doherty, 1886: 110. Type Genus: Gerydus Boisduval, 1836 [= Miletus Hubner, 1819].

Corbet, A. S., 1939a. A revision of the Malayan species of Allotinus Felder & Felder (Lepidoptera: Lycaenidae) Trans. R. ent. Soc. Lond. 89: 63–78, 1pl., 21figs. Reference page.
Doherty, W., 1886. A list of butterflies taken in Kumaon. J. Asiat. Soc. Beng. 55(2): 103–140. BHL. Reference page.
Eliot, J.N. 1973. The higher classification of the Lycaenidae (Lepidoptera): a tentative arrangement. Bulletin of the British Museum (Natural History), entomology 28(6): 371-505. Reference page.
Eliot, J.N. 1986. A review of the Miletini (Lepidoptera: Lycaenidae). Bull. Br. Mus. nat. Hist. (Ent.) 53(1): 1–105. Reference page.
Espeland, M., Breinholt, J.W., Willmott, K.R., Warren, A.D., Vila, R., Toussaint, E.F.A., Maunsell, S.C., Aduse-Poku, K., Talavera, G., Eastwood, R., Jarzyna, M.A., Guralnick, R., Lohman, D.J., Pierce, N.E. & Kawahara, A.Y. 2018. A Comprehensive and Dated Phylogenomic Analysis of Butterflies. Current Biology 28(5): 770–778. DOI: 10.1016/j.cub.2018.01.061 Reference page.
Hübner, J. 1816–1826. Verzeichniss bekannter Schmettlinge. Augsburg. Papilionoidea. BHL Reference page.
Kaliszewska, Z.A., Lohman, D.J., Sommer, K., Adelson, G., Rand, D.B., Mathew, J., Talavera, G. & Pierce, N.E., 2015. When caterpillars attack: Biogeography and life history evolution of the Miletinae (Lepidoptera: Lycaenidae). Evolution 69(3): 571–588. DOI: 10.1111/evo.12599 Paywall. Reference page.


Williams, Mark C., 2008: Checklist of Afrotropical Papilionoidea and Hesperioidea

Vernacular names
日本語: カニアシシジミ亜科
中文: 云灰蝶亚科

Miletinae is a subfamily of the family Lycaenidae of butterflies, commonly called harvesters and woolly legs, and virtually unique among butterflies in having predatory larvae. Miletinae are entirely aphytophagous (do not feed on plants). The ecology of the Miletinae is little understood, but adults and larvae live in association with ants, and most known species feed on Hemiptera (aphids, coccids, membracids, and psyllids), though some, like Liphyra, feed on the ants themselves. The butterflies, ants, and hemipterans, in some cases, seem to have complex symbiotic relationships benefiting all.[1]

Tribe Miletini
Allotinus C. & R. Felder, [1865] — Indomalayan realm
Lontalius Eliot, 1986 — Indomalayan realm
Miletus Hübner, [1819] — Indomalayan realm
Logania Distant, 1884 — Indomalayan realm
Megalopalpus Röber, [1886] — Afrotropical realm
Tribe Spalgini
Spalgis Moore, 1879 — Indomalayan realm, Afrotropical realm
Feniseca Grote, 1869 — Nearctic realm (one species, Feniseca tarquinius)
Taraka Doherty, 1889 — East Asian Palearctic realm, Indomalayan realm (sometimes in a separate tribe: Tarakini)
Tennenta Müller, 2017 — Australasian realm
Tribe Lachnocnemini
Lachnocnema Trimen, 1887 — Afrotropical realm
Thestor Hübner, [1819] — Afrotropical realm
Tribe Liphyrini (formerly a separate subfamily: Liphyrinae)
Euliphyra Holland, 1890 — Afrotropical realm
Aslauga Kirby, 1890 — Afrotropical realm
Liphyra Westwood, [1864] — Indomalayan realm, Australasian realm


Lohman, D.J.; Samarita, V.U. 2009: The biology of carnivorous butterfly larvae (Lepidoptera: Lycaenidae: Miletinae: Miletini) and their ant-tended hemipteran prey in Thailand and the Philippines. Journal of natural history, 43: 569-581.

Bernard d'Abrera (1986) Butterflies of the Oriental Region. Part 3: Lycaenidae and Riodinidae Hill House Publishers ISBN 0-9593639-4-7
Bernard d'Abrera, (1980) Butterflies of the Afrotropical region based on Synonymic catalogue of the butterflies of the Ethiopian region by R.H. Carcasson. Lansdown Editions in association with E.W. Classey, Melbourne ISBN 0-7018-1029-7
Kaliszewska, Z.A., Lohman, D.J., Sommer, K., Adelson, G., Rand, D.B., Mathew, J., Talavera, G. & Pierce, N.E. 2015. When caterpillars attack: Biogeography and life history evolution of the Miletinae (Lepidoptera: Lycaenidae). Evolution 69(3): 571–588.

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