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Superregnum: Eukaryota
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Endopterygota
Superordo: Panorpida
Cladus: Amphiesmenoptera
Ordo: Lepidoptera
Subordo: Glossata
Cladus: Coelolepida
Cladus: Myoglossata
Cladus: Neolepidoptera
Infraordo: Heteroneura
Cladus: Eulepidoptera
Cladus: Ditrysia
Cladus: Apoditrysia
Cladus: Obtectomera
Superfamilia: Papilionoidea

Familia: Lycaenidae
Subfamilia: Theclinae
Tribus: Luciini
Genus: Philiris
Species: (70)
P. agatha – P. albicostalis – P. albihumerata – P. albiplaga – P. amethysta – P. angabunga – P. apicalis – P. aquamarina – P. argenteus – P. azula – P. baiteta – P. biplaga – P. bubalisatina – P. cadmica – P. caelestis – P. diana – P. dinawa – P. doreia – P. elegans – P. fulgens – P. gloriosa – P. harterti – P. helena – P. hemileuca – P. hindenburgensis – P. hypoxantha – P. ianthina – P. ignobilis – P. ilias – P. innotatus – P. intensa – P. kumusiensis – P. lavendula – P. lucescens – P. lucina – P. maculata – P. marginata – P. mayri – P. melanacra – P. misimensis – P. moira – P. moluccana – P. montigena – P. nitens – P. oreas – P. pagwi – P. papuanus – P. parsonsi – P. petriei – P. phengotes – P. philotas – P. philotoides – P. praeclara – P. putih – P. radicala – P. refusa – P. remissa – P. sappheira – P. satis – P. scintillata – P. siassi – P. sibatanii – P. subovata – P. tapini – P. tombara – P. unipunctata – P. vicina – P. violetta – P. zadne – P. ziska

Philiris Röber, 1891: 317.

Type species: Thecla ilias Felder, 1860: 454, by monotypy.

Philiris Röber, 1891: 317.
Philiris Röber, in Staudinger & Schatz, 1892: 273.


Bethune-Baker, G. T., 1904. New Lepidoptera from British New Guinea. Novit. zool. 11:367-429, 3 pls. Archive plates. Reference page.
Butler, A. G., 1876. On a collection of Lepidoptera from Port Moresby. Annals and Magazine of natural History 4(18): 240–249. Archive. Reference page.
Cassidy, A.C., 2003. New information on lycaenid butterflies of the South-East Asian Islands. Transactions of the Lepidopterological Society, Japan 54(2): 73–82. pdf. Reference page.
Felder, C. 1860. Lepidopterorum Amboinensium species novae diagnosibus. S-B. Akad. Wiss. Wien 40(11): 447–468. BHL Reference page.
Miskin, W.H. 1874. New Lycaenidae from Queensland. Ent. mon. Mag. 11: 165. Reference page.
Müller, C. J., 2002. New butterfly taxa from New Ireland, Papua New Guinea (Lepidoptera: Nymphalidae and Lycaenidae). Australian Entomologist 29 (1): 29–36. Reference page.
Müller, C.J. 2014: Six new species of Philiris Röber, 1891 (Lepidoptera, Lycaenidae) from Papua New Guinea. ZooKeys 395: 33–55. DOI: 10.3897/zookeys.395.7110 Reference page.
Parsons, M.J. 1998. The Butterflies of Papua New Guinea xvi+736pp., 136 pls. Academic Press, London, ISBN 0125455550. Reference page.
Ring, L. R. & Olive, J. C., 1997. The specific status of Philiris sappheira Sands (Lepidoptera, Lycaenidae), with descriptions of a new subspecies from Australia. Australian Entomologist 24 (2): 65–72.
Röber, J., 1891. Beitrag zur kenntniss der Indo-Australischen lepidopteren fauna. Tijdschr. Ent. 34: 261-334. Reference page.
Sands, D.P.A. 1979. New species of Philiris Röber (Lepidoptera: Lycaenidae) from Papua New Guinea. Journal of the Australian Entomological Society 18(2): 127-133. PDF. Reference page.
Sands, D.P.A., 1981b. Biogeography and speciation in the Lycaenid butterfly tribe Luciini. News Bull. Ent. Soc. Qd. 9:18-28. Reference page.
Sands, D.P.A., 1981. The ecology, biogeography and systematics of the tribe Luciini. PhD thesis, University of Queensland, Australia. 2 vols: 1: 1-584 (text); 2: 1-168 (2pls & figs). (Unpublished). DOI: 10.14264/uql.2014.573 Reference page.
Staudinger, O., 1892, in Staudinger, O. & Schatz, E., 1884–1892. Exotische Schmetterlinge 2. Die Familien und Gattungen der Tagfalter 284pp, 100 pls. BHL text, plates. Reference page.
Tite, G.E. 1963. A revision of the genus Candalides and allied genera (Lepidoptera: Lycaenidae). Bulletin of the British Museum (Natural History) 14: 197–259, 4 pls, 119 text figs. BHL Reference page.

Philiris is a genus of butterflies in the family Lycaenidae. The species of this genus are found in the Australasian realm (New Guinea, Bismarck Islands, Molucca Islands and eastern Australia). Philiris was erected by Julius Röber in 1891. It is a speciose genus; Tite decided on 56 species,[1] Sands[2][3] added 11 species and placed the taxa into 21 species groups (broadly accepted by Parsons [4] Most species are on New Guinea.Tite considered Philiris and Candalides Hübner, 1819 to be sisters. Eliot (1973)[5] placed Philiris in Luciini Waterhouse & Lyell, 1914,

Philiris agatha
Philiris albicostalis
Philiris albihumerata
Philiris albiplaga
Philiris amethysta
Philiris angabunga
Philiris apicalis
Philiris aquamarina
Philiris argenteus
Philiris azula
Philiris baiteta Müller, 2014[6]: 41–44 
Philiris biplaga
Philiris bubalisatina Müller, 2014[6]: 39–41 
Philiris cadmica
Philiris caelestis
Philiris cyana
Philiris diana
Philiris dinawa
Philiris doreia
Philiris elegans
Philiris fulgens
Philiris gloriosa
Philiris harterti
Philiris helena
Philiris hemileuca
Philiris hindenburgensis Müller, 2014[6]: 47–49 
Philiris hypoxantha
Philiris ianthina
Philiris ignobilis
Philiris ilias
Philiris innotatus
Philiris intensa
Philiris kumusiensis
Philiris lavendula
Philiris lucescens
Philiris maculata
Philiris marginata
Philiris mayri
Philiris melanacra
Philiris misimensis
Philiris moira
Philiris moluccana
Philiris montigena
Philiris nitens
Philiris oreas
Philiris pagwi
Philiris parsonsi Müller, 2014[6]: 49–53 
Philiris petriei Müller, 2014[6]: 35–39 
Philiris phengotes
Philiris philotas
Philiris philotoides
Philiris praeclara
Philiris putih
Philiris radicala
Philiris refusa Müller, 2014[6]: 44–47 
Philiris remissa
Philiris satis
Philiris scintillata
Philiris siassi
Philiris sibatanii
Philiris subovata
Philiris tapini
Philiris tombara
Philiris unipunctata
Philiris vicina
Philiris violetta
Philiris zadne
Philiris ziska


Tite, 1963 A revision of the genus Candalides and allied genera (Lepidoptera: Lycaenidae) Bull. Br. Mus. (Nat. Hist.) (Entomol.) 14 (5) : 197-259, pl. 1-4
Sands, 1979 New species of Philiris Röber (Lepidoptera: Lycaenidae) from Papua New Guinea J. Aust. Entomol. Soc. 18 (2): 127-133
Sands, 1981 New species of Philiris Röber (Lepidoptera: Lycaenidae) from mainland New Guinea J. Aust. Entomol. Soc. 20: 89–96
Parsons, M. J. 1998 The butterflies of Papua New Guinea. Their systematics and biology. — London (Academic Press), 737 pp., 26 + 139 pls.)
Eliot, J. N. 1973. More Revisional Notes on Oriental Butterflies. Entomologist 106: 1–15, 10 figs.
Müller, Chris J. (2014). "Six new species of Philiris Röber, 1891 (Lepidoptera, Lycaenidae) from Papua New Guinea". ZooKeys (395): 33–55. doi:10.3897/zookeys.395.7110. PMC 3978236. PMID 24715788.

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