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RatardinaeSuperregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Endopterygota
Superordo: Panorpida
Cladus: Amphiesmenoptera
Ordo: Lepidoptera
Subordo: Glossata
Cladus: Coelolepida
Cladus: Myoglossata
Cladus: Neolepidoptera
Infraordo: Heteroneura
Cladus: Eulepidoptera
Cladus: Ditrysia
Cladus: Apoditrysia
Superfamilia: Cossoidea

Familia: Cossidae
Subfamily: Ratardinae
Genera: Callosiope - Ratarda - Shisa - Sumatratarda

The Ratardinae are a small subfamily of large moths from Southeast Asia.

Taxonomy and systematics

Ratardinae is a small subfamily of moths formerly placed in its own family Ratardidae and related to (and often included within) the Cossidae. Three genera are known, one quite recently described.[1] One species, "Shisa" excellens, was originally placed in the Lymantriidae.[2] The moths are large with rounded wings and strongly spotted wing patterns, and "pectinate" antennae. The relationships of this group to other Cossoidea needs reassessment, once suitable samples are available, with molecular data.[3]

About 13 relictually distributed species are restricted to Southeast Asia, occurring in Borneo, Sumatra, Peninsular Malaysia, northeastern Himalayas, and Taiwan.[4]

The adults are very rarely found, feeble-flying, and occasionally are attracted to light, but more likely to be found flying by day.[5]


These large moths are so incredibly rarely found and their habitats under such massive threat from large-scale conversion of rainforest in Southeast Asia that their conservation status should be seriously considered and dedicated surveys conducted to assess their distribution and biology. One species (Ratarda melanoxantha) is probably protected by virtue of its occurrence in Mount Kinabalu National Park on Borneo, where it was found once.[6]

(Kobes and Ronkay, 1990).
(Owada, 1993; Holloway, 1998: 9). Archived from the original on September 16, 2007. Retrieved March 17, 2007. {{cite web}}: Missing or empty |title= (help)
Holloway, 1986: 41-42; Heppner and Wang, 1987; Heynderycx, 2003
(Holloway, 1986: 41-42).

(Holloway, 1986).

Edwards, E.D., Gentili, P., Horak, M., Kristensen, N.P. and Nielsen, E.S. (1999). The cossoid/sesioid assemblage. Ch. 11, pp. 181–195 in Kristensen, N.P. (Ed.). Lepidoptera, Moths and Butterflies. Volume 1: Evolution, Systematics, and Biogeography. Handbuch der Zoologie. Eine Naturgeschichte der Stämme des Tierreiches / Handbook of Zoology. A Natural History of the phyla of the Animal Kingdom. Band / Volume IV Arthropoda: Insecta Teilband / Part 35: 491 pp. Walter de Gruyter, Berlin, New York.
Heynderycx, J. 2003. Les Ratardinae. Lambillionea, 103(1): 133-134.
Heppner J.B. and Wang, H.Y. 1987. A rare moth, Ratarda tertia Strand (Lepidoptera: Ratardidae), from Palin Taiwan. Táiwa-n she(nglì bówùgua(n bànniánka-n (Táiwa-n she(nglì bówùgua(n bànniánka-n), 40: 91-94.
Holloway, J.D. (1986). The Moths of Borneo: Key to Families: Families Cossiae, Metarbelidae, Ratardidae, Dudgeonidae, Epipyropidae and Limacodidae. Malayan Nauture Journal, 40: 1-166.
Holloway, J.D. (1998). The Moths of Borneo: Families Castniidae, Callidulidae, Drepanidae and Uraniidae. Malayan Nauture Journal, 52: 1-155.
Kobes, L.W.R. and Ronkay, L. (1990). The Ratardidae of Sumatra, Heterocera Sumatrana, 6: 79-100.
Owada, M. (1993). The systematic position of Shisa excellens (Lepidoptera and Ratardidae). Japanese Journal of Entomology, 61(2): 251-260.

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