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Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Crustacea
Superclassis: Multicrustacea
Classis: Malacostraca
Subclassis: Eumalacostraca
Superordo: Eucarida
Ordo: Decapoda
Subordo: Pleocyemata
Infraordo: Brachyura
Sectio: Eubrachyura
Subsectio: Thoracotremata
Superfamilia: Ocypodoidea

Familia: Ocypodidae
Subfamiliae: Gelasiminae – Heloeciinae - Macrophthalminae - Ocypodinae

Overview of genera

Gelasimus – Tubuca – ...

Ocypode - Uca -


Ocypodidae Rafinesque, 1815

Rafinesque, C.S. 1815. Analyse de la nature, ou tableau de l'univers et des corps organisés. Palerme: L'Imprimerie de Jean Barravecchia. 224 pp. BHL Reference page.
Masunari, S., Martins, S.B. & Anacleto, A.F.M. 2020. An illustrated key to the fiddler crabs (Crustacea, Decapoda, Ocypodidae) from the Atlantic coast of Brazil. ZooKeys, 943: 1–20. DOI: 10.3897/zookeys.943.52773 Open access Reference page.
Shih, H-T., Chan, B.K-K. & Ng, P.K.L. 2018. Tubuca alcocki, a new pseudocryptic species of fiddler crab from the Indian Ocean, sister to the southeastern African T. urvillei (H. Milne Edwards, 1852) (Crustacea, Decapoda, Brachyura, Ocypodidae). ZooKeys 747: 41-62. DOI: 10.3897/zookeys.747.23468. Reference page.

Vernacular names
日本語: スナガニ科

The Ocypodidae are a family of semiterrestrial crabs that includes the ghost crabs and fiddler crabs. They are found on tropical and temperate shorelines around the world.

Some genera previously included in the family are now treated as members of separate families in the superfamily Ocypodoidea, such as the Dotillidae and Macrophthalmidae. In 2016, the genus Uca was split into 13 genera by elevating its subgenera to genus rank, among other things.[1] Further refinements have been made to the organization of the family, which has 4 subfamilies, about 13 extant genera, and about 180 described species as of 2020.[2][3][4]

The majority of the Ocypodidae species are fiddler crabs, the exceptions being the members of the genus Ocypode, ghost crabs, and the genus Ucides, mangrove crabs.[2][3]

These genera belong to the family Ocypodidae:[2][5][6]

Afruca Crane, 1975
Austruca Bott, 1973
Cranuca Beinlich & von Hagen, 2006
Gelasimus Stimpson 1862
Leptuca Bott, 1973
Minuca Bott, 1954
Ocypode Weber, 1795 - (ghost crabs)
Paraleptuca Bott, 1973
Petruca N.g.Shih & Christy, 2015
Tubuca Bott, 1973
Uca Leach, 1814
Ucides Rathbun, 1897 - (mangrove crabs)
Xeruca Shih, 2015
† Chondromaia Feldmann, et al., 2013
† Eoinachoides Van Straelan, 1933
† Micromaia Bittner, 1875
† Mithracia Bell, 1858
† Nanomaja Müller & Collins, 1991
† Panticarcinus Collins & Saward, 2006
† Periacanthus Bittner, 1875
† Pisomaja Lorenthey, 1929
† Wilsonimaia Blow & Manning, 1996


Shih, Hsi-Te; Ng, Peter K. L.; Davie, Peter J. F.; Schubart, Christoph D.; et al. (2016). "Systematics of the family Ocypodidae Rafinesque, 1815 (Crustacea: Brachyura), based on phylogenetic relationships, with a reorganization of subfamily rankings and a review of the taxonomic status of Uca Leach, 1814, sensu lato and its subgenera". The Raffles Bulletin of Zoology. 64.
Rosenberg, Michael S. (2019). "A fresh look at the biodiversity lexicon for fiddler crabs (Decapoda: Brachyura: Ocypodidae). Part 1: Taxonomy". Journal of Crustacean Biology. 39 (6).
"WoRMS taxon details, Ocypodidae Rafinesque, 1815". World Register of Marine Species. Retrieved 2020-11-15.
"Ocypodidae". GBIF. Retrieved 2020-11-15.
Sammy De Grave; N. Dean Pentcheff; Shane T. Ahyong; et al. (2009). "A classification of living and fossil genera of decapod crustaceans" (PDF). Raffles Bulletin of Zoology. Suppl. 21: 1–109.

"Family Ocypodidae Rafinesque 1815 (crab)". Paleo DB, Paleobiology Database. Retrieved 2020-11-15.

Further reading

Stebbing, Thomas Roscoe Rede (1893). A History of Crustacea: Recent Malacostraca. D. Appleton.


Biology Encyclopedia

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