Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: ParaHoxozoa
Cladus: Bilateria
Cladus: Nephrozoa
Superphylum: Deuterostomia
Phylum: Chordata
Cladus: Olfactores
Subphylum: Vertebrata
Infraphylum: Gnathostomata
Megaclassis: Osteichthyes
Cladus: Sarcopterygii
Cladus: Rhipidistia
Cladus: Tetrapodomorpha
Cladus: Eotetrapodiformes
Cladus: Elpistostegalia
Superclassis: Tetrapoda
Cladus: Batrachomorpha
Classis: Amphibia
Subclassis: Lissamphibia
Superordo: Batrachia
Cladus: Salientia
Ordo: Anura
Familia: Megophryidae
Subfamilia: Megophryinae
Genus: Pelobatrachus
Species (7): P. baluensis – P. edwardinae – P. kalimantanensis – P. kobayashii – P. ligayae – P. nasutus – P. stejnegeri
Name
Pelobatrachus Beddard, 1908: 909
Type species: Ceratophryne nasuta Schlegel, 1858, by monotypy.
Synonyms
Ceratophryne Schlegel, 1858: 56 [synonymised with Megophrys by Inger (1954: 223), and with Pelobatrachus by Lyu et al. (2021: 39)]
Type species: Ceratophryne nasuta Schlegel, 1858, by subsequent designation of Gorham (1966: 15).
Borneophrys Delorme, Dubois, Grosjean & Ohler, 2006: 15 [synonymised with Megophrys by Chen et al. (2017: 41), and with Pelobatrachus by Lyu et al. (2021: 39)]
Type species: Megophrys edwardinae Inger, 1989, by original designation.
References
Primary references
Beddard, F.E. [1907] 1908. Contributions to the knowledge of the anatomy of the batrachian family Pelobatidae. Proceedings of the Zoological Society of London 1907: 871–911.
Delorme, M., Dubois, A., Grosjean, S. & Ohler, A. 2006. Une nouvelle ergotaxinomie des Megophryidae (Amphibia, Anura). Alytes 24(1-4): 6–21. Reference page.
Chen, J.-M., Zhou, W.-W., Poyarkov, Jr., N.A., Stuart, B.L., Brown, R.M., Lathrop, A., Wang, Y.-Y., Yuan, Z.-Y., Jiang, K., Hou, M., Chen, H.-M., Suwannapoom, C., Nguyen, S.N., Duong, T.V., Papenfuss, T.J., Murphy, R.W., Zhang, Y.-P. & Che, J. 2017. A novel multilocus phylogenetic estimation reveals unrecognized diversity in Asian horned toads, genus Megophrys sensu lato (Anura: Megophryidae). Molecular Phylogenetics and Evolution 106: 28–43. DOI: 10.1016/j.ympev.2016.09.004 Paywall Reference page.
Lyu, Z.-T., Zeng, Z.-C., Wang, J., Liu, Z.-Y., Huang, Y.-Q., Li, W.-Z. & Wang, Y.-Y. 2021. Four new species of Panophrys (Anura, Megophryidae) from eastern China, with discussion on the recognition of Panophrys as a distinct genus. Zootaxa 4927(1): 9–40. DOI: 10.11646/zootaxa.4927.1.2 Paywall Reference page.
Additional references
Lyu, Z.-T., Qi, S., Wang, J., Zhang, S.-Y., Zhao, J., Zeng, Z.-C., Wan, H., Yang, J.-H., Mo, Y.-M. & Wang, Y.-Y. 2023. Generic classification of Asian horned toads (Anura: Megophryidae: Megophryinae) and monograph of Chinese species. Zoological Research 44(2): 380–450. DOI: 10.24272/j.issn.2095-8137.2022.372 Open access Reference page.
Vernacular names
English: Clay Horned Toads
Pelobatrachus is a genus of frogs in the family Megophryidae. It was formerly synonymized with Megophrys until 2021, when it was revived as a distinct genus. They inhabit Southeast Asia, namely the Malay Peninsula, Sumatra, Borneo and the Philippines. Their common name is clay horned toads.[1][2][3]
Taxonomy
The following species are recognized in the genus Pelobatrachus:
Pelobatrachus baluensis (Boulenger, 1899) — Kinabalu horned frog
Pelobatrachus edwardinae (Inger, 1989) — Edwardina's horned frog
Pelobatrachus kalimantanensis (Munir, Hamidy, Matsui, Iskandar, Sidik, and Shimada, 2019) — Kalimantan horned frog
Pelobatrachus kobayashii (Malkmus and Matsui, 1997) — Kobayashi's horned frog
Pelobatrachus ligayae (Taylor, 1920) — Palawan horned frog
Pelobatrachus nasutus (Schlegel, 1858) — long-nosed horned frog or Malayan horned frog
Pelobatrachus stejnegeri (Taylor, 1920) — Mindanao horned frog
References
"Pelobatrachus Beddard, 1908 "1907" | Amphibian Species of the World". amphibiansoftheworld.amnh.org. Retrieved 2022-09-03.
"AmphibiaWeb - Megophryidae". amphibiaweb.org. Retrieved 2022-09-03.
Lyu, Zhi-Tong; Zeng, Zhao-Chi; Wang, Jian; Liu, Zu-Yao; Huang, Ya-Qiong; Li, Wen-Zhou; Wang, Ying-Yong (2021-02-11). "Four new species of Panophrys (Anura, Megophryidae) from eastern China, with discussion on the recognition of Panophrys as a distinct genus". Zootaxa. 4927 (1): 9–40. doi:10.11646/zootaxa.4927.1.2. ISSN 1175-5334.
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