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Branta bernicla

Branta bernicla (*)

Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Superphylum: Deuterostomia
Phylum: Chordata
Subphylum: Vertebrata
Infraphylum: Gnathostomata
Megaclassis: Osteichthyes
Cladus: Sarcopterygii
Cladus: Rhipidistia
Cladus: Tetrapodomorpha
Cladus: Eotetrapodiformes
Cladus: Elpistostegalia
Superclassis: Tetrapoda
Cladus: Reptiliomorpha
Cladus: Amniota
Classis: Reptilia
Cladus: Eureptilia
Cladus: Romeriida
Subclassis: Diapsida
Cladus: Sauria
Infraclassis: Archosauromorpha
Cladus: Crurotarsi
Divisio: Archosauria
Cladus: Avemetatarsalia
Cladus: Ornithodira
Subtaxon: Dinosauromorpha
Cladus: Dinosauriformes
Cladus: Dracohors
Cladus: Dinosauria
Ordo: Saurischia
Cladus: Eusaurischia
Subordo: Theropoda
Cladus: Neotheropoda
Cladus: Averostra
Cladus: Tetanurae
Cladus: Avetheropoda
Cladus: Coelurosauria
Cladus: Tyrannoraptora
Cladus: Maniraptoromorpha
Cladus: Maniraptoriformes
Cladus: Maniraptora
Cladus: Pennaraptora
Cladus: Paraves
Cladus: Eumaniraptora
Cladus: Avialae
Infraclassis: Aves
Cladus: Euavialae
Cladus: Avebrevicauda
Cladus: Pygostylia
Cladus: Ornithothoraces
Cladus: Ornithuromorpha
Cladus: Carinatae
Parvclassis: Neornithes
Cohors: Neognathae
Cladus: Pangalloanserae
Cladus: Galloanseres
Ordo: Anseriformes

Familia: Anatidae
Subfamilia: Anserinae
Genus: Branta
Species: Branta bernicla
Subspecies: B. b. bernicla – B. b. hrota – B. b. nigricans – B. b. orientalis

Branta bernicla (Linnaeus, 1758)

Anas bernicla (protonym)


Linnaeus, C. 1758. Systema Naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Editio Decima, Reformata. Tomus I. Holmiæ (Stockholm): impensis direct. Laurentii Salvii. 824 pp. DOI: 10.5962/bhl.title.542 BHL p. 124 BHL Reference page.
Buckley, P.A., & Mitra, S.S. 2002. Three geese resembling "Gray-bellied Brant" / "Lawrence's Brant" from Long Island, New York. North American Birds 56(4): 502–507. PDF.
Reed, A., Ward, D.H., Derksen, D.V. & Sedinger, J.S. 2013. Brant Branta bernicla. The Birds of North America, Cornell Lab of Ornithology. DOI: 10.2173/bna.337

Vernacular names
беларуская: Казарка чорная
български: Черна гъска
brezhoneg: Garreli du
català: Oca de collar
čeština: Berneška tmavá
Cymraeg: Gŵydd Ddu
dansk: Knortegås
Deutsch: Ringelgans
Ελληνικά: Δαχτυλιδόχηνα
English: Brant Goose
Esperanto: Kolringa berniklo
español: Barnacla carinegra
eesti: Mustlagle
euskara: Branta musubeltz
suomi: Sepelhanhi
føroyskt: Helsigás
Nordfriisk: Groltergus
français: Bernache cravant
Frysk: Swartbúkrotgoes
Gaeilge: Cadhan
Gàidhlig: Gèadh Got
galego: Ganso cabecinegro
Gaelg: Guiy breck
עברית: ברנטה כהה
hrvatski: Grivasta guska
magyar: Örvös lúd
íslenska: Margæs
italiano: Oca colombaccio
ᐃᓄᒃᑎᑐᑦ/inuktitut: Nerlernaq
日本語: コクガン
қазақша: Қарашақаз
한국어: 흑기러기
Limburgs: Rotgajs
lietuvių: Paprastoji berniklė
latviešu: Melngalvas tumšzoss
монгол: Хар алаг галуу
Nederlands: Rotgans
norsk nynorsk: Ringgås
norsk: Ringgås
polski: Bernikla obrożna
português: Ganso-de-faces-negras
rumantsch: Auca dal cularin
русский: Чёрная казарка
саха тыла: Хардырҕас
davvisámegiella: Gierdočuonjá
slovenčina: Bernikla tmavá
slovenščina: Crna gos
српски / srpski: Гриваста гуска, Grivasta guska
svenska: Prutgås
Türkçe: Yosun kazı
українська: Чорна казарка
Zeêuws: Rothanze
中文: 黑雁

The brant or brent goose (Branta bernicla) is a small goose of the genus Branta. There are three subspecies, all of which winter along temperate-zone sea-coasts and breed on the high-Arctic tundra.

The Brent oilfield, was named after the species.[3]

Wintering at the Wadden Sea, Germany
Feeding in Long Island Sound, Connecticut
Eggs, Collection Museum Wiesbaden

The brant is a small goose with a short, stubby bill. It measures 55–66 cm (22–26 in) long, 106–121 cm (42–48 in) across the wings and weighs 0.88–2.2 kg (1.9–4.9 lb).[4][5][6][7] The under-tail is pure white, and the tail black and very short (the shortest of any goose).

The species is divided into three subspecies:[8]

Dark-bellied brant goose B. b. bernicla (Linnaeus, 1758)
Pale-bellied brant goose B. b. hrota (Müller, 1776) (also known as light-bellied brent goose in Europe, and Atlantic brant in North America)
Black brant goose B. b. nigricans (Lawrence, 1846) (sometimes also known as the Pacific brant in North America)

Some DNA evidence suggests that these forms are genetically distinct; while a split into three separate species has been proposed, it is not widely accepted, with other evidence upholding their maintenance as a single species.

The body of the dark-bellied form B. b. bernicla is fairly uniformly dark grey-brown all over, the flanks and belly not significantly paler than the back. The head and neck are black, with a small white patch on either side of the neck. With a population of about 250000, it breeds on the Arctic coasts of central and western Siberia and winters in western Europe, with over half the population in southern England, the rest between northern Germany and north-western France.

The pale-bellied brant B. b. hrota appears blackish-brown and light grey in colour. The body is different shades of grey-brown all over, the flanks and belly are significantly paler than the back and present a marked contrast. The head and neck are black, with a small white patch on either side of the neck. The total population is about 250000, with the main population breeding in northeastern Canada and wintering along the Atlantic coast of the U.S. from Maine to Georgia,[4] and two smaller populations, one breeding in Franz Josef Land, Svalbard, and northeastern Greenland and wintering in Denmark, northeast England, and Scotland, and the other breeding in the far-northeastern Canadian islands and wintering in Ireland, southwest England, and in a small but significant area, le Havre de Regnéville, centered on the Sienne Estuary in Manche (Northern France).[9] In Ireland it is recorded in winter from a number of areas including Lough Foyle, Strangford Lough, Tralee Bay and Castlemaine Harbour.[10][11][12]

The black brant B. b. nigricans appears blackish-brown and white in colour. This form is a very contrastingly black and white bird, with a uniformly dark sooty-brown back, similarly coloured underparts (with the dark colour extending furthest back of the three forms) and a prominent white flank patch; it also has larger white neck patches, forming a near-complete collar. The population of about 125000 breeds in northwestern Canada, Alaska and eastern Siberia, and winters mostly on the west coast of North America from southern Alaska to California, but also some in east Asia, mainly Japan, also Korea and China. The population has been as high as 200,000 in 1981, and as low as 100,000 in 1987.[13]

The Asian populations of the black brant populations had previously been regarded as a separate subspecies B. b. orientalis based on purported paler upperparts coloration; however, it is generally now believed that this is not correct, and they are assigned to B. b. nigricans.

A fourth form (known variously as gray brant, intermediate brant, or grey-bellied brent goose) has been proposed, although no formal subspecies description has been made as yet, for a population of birds breeding in central Arctic Canada (mainly Melville Island), and wintering on Puget Sound on the American west coast around the U.S./Canada border. These birds are intermediate in appearance between black brant and pale-bellied brant, having brown upperparts and grey underparts which give less of a contrast with the white flank patch. It has also been proposed that, rather than being a separate subspecies, it is actually a result of interbreeding between these two forms, given that this population exhibits mixed characters.

Individual birds when wintering generally remain in loose family-groups, together with others of the same sub-species, but there is overlap in some areas (for example Western Europe, see above); and this is also true in the breeding colonies. Outside the breeding season, individuals with characteristics of any subspecies may occasionally turn up with regular migrants, and there has been debate as to whether this is related to migration routing accidents, or to breeding range overlap, or even interbreeding.
In a defensive position
In flight

The brant goose was strictly coastal bird in winter, rarely leaving tidal estuaries, where it feeds on eelgrass (Zostera marina)[8] and the seaweed, sea lettuce (Ulva).[4][14] On the east coast of North America, the inclusion of sea lettuce is a recent change to their diet, brought about by a blight on eelgrass in 1931. This resulted in the near-extirpation of the brant. The few that survived changed their diet to include sea lettuce until the eelgrass eventually began to return. Brants have maintained this diet ever since as a survival strategy.[15] A similar collapse in eelgrass in Ireland in the 1930s also negatively impacted the population.[16] In recent decades, it has started using agricultural land a short distance inland, feeding extensively on grass and winter-sown cereals. It has been suggested that they learnt this behaviour by following other species of geese. Food resource pressure may also be important in forcing this change, as the world population increased over 10-fold to 400,000-500,000 by the mid-1980s, possibly reaching the carrying capacity of the estuaries. In the breeding season, it uses low-lying wet coastal tundra for both breeding and feeding. The nest is bowl-shaped, lined with grass and down, in an elevated location, often near a small pond.

The brant goose is one of the species to which the Agreement on the Conservation of African-Eurasian Migratory Waterbirds applies.

Branta is a Latinised form of Old Norse brandgás, "burnt (black) goose", and bernicla is the medieval Latin name for the barnacle.

The brant and the similar barnacle goose were previously considered one species, formerly believed to be the same creature as the crustacean.[17] That myth can be dated back to at least the 12th century. Gerald of Wales claimed to have seen these birds hanging down from pieces of timber, William Turner accepted the theory, and John Gerard claimed to have seen the birds emerging from their shells. This myth arose because in the 1100's the migration of birds was unknown, but it was known that none of these birds was ever seen nesting, nor were eggs found, nor were goslings seen.[18]

The legend persisted until the end of the 18th century. In County Kerry, until relatively recently, Catholics could eat this bird on a Friday because it counted as fish.[19]

BirdLife International (2016). "Branta bernicla". IUCN Red List of Threatened Species. 2016: e.T22679946A85966135. doi:10.2305/IUCN.UK.2016-3.RLTS.T22679946A85966135.en. Retrieved 12 November 2021.
Lewis, Tyler L.; Ward, David H.; Sedinger, James S.; Reed, Austin; Derksen, Dirk V.; Carboneras, Carles; Christie, David; Kirwan, Guy M. (4 March 2020). "Brant (Branta bernicla)". Birds of the World. doi:10.2173/bow.brant.01.
"UK Brent Oil". One Financial Markets. Archived from the original on 15 February 2016. Retrieved 25 August 2014.
"Norsk Polarinstitutt". Norsk Polarinstitutt. Retrieved 17 September 2020.
"Brant". All About Birds. Cornell Lab of Ornithology. Retrieved 18 October 2011.
"Brent Goose". Retrieved 18 October 2011.
Dunning, John Barnard (2007). CRC handbook of avian body masses (2nd ed.). Boca Raton: CRC Press. p. 38. ISBN 978-1-4200-6445-2. OCLC 310154556.
Ganter, Barbara (2000). "Seagrass (Zostera spp.) as food for brent geese (Branta bernicla): an overview" (PDF). Helgoland Marine Research (54): 63–70. doi:10.1007/s101520050003. Retrieved 8 December 2021. (subspecies, breeding/wintering ranges with flyway maps, diet)
Maheo, Roger (1976). "The Brent Geese of France, with special reference to the Golfe du Morbihan". Wildfowl. 27: 45–53.
Ruttledge, R.F. 1966 Ireland's Birds their Distribution and Migration. H.F. & G. Witherby Ltd
"Brent Goose (Light-bellied)". BirdWatch Ireland. Retrieved 17 September 2020.
"Branta bernicla hrota Pale-bellied Brent Goose". Retrieved 17 September 2020.
Slimbridge: Black Brant[permanent dead link]
"Brant". Audubon. 13 November 2014. Retrieved 17 September 2020.
Book of North American Birds. 1990.
"Where wild geese fly". The Irish Times. 17 August 2002. Retrieved 17 September 2020.
Jobling, James A (2010). The Helm Dictionary of Scientific Bird Names. London: Christopher Helm. pp. 70, 77. ISBN 978-1-4081-2501-4.
Goose Barnacles: Undulating Creatures Retrieved 2011-11-28.

Cocker, Mark; Mabey, Richard (2005). Birds Britannica. London: Chatto & Windus. pp. 78–80. ISBN 0-7011-6907-9.

Further reading
Millington, Richard (1997). Separation of Black Brant, Dark-bellied Brent Goose and Pale-bellied Brent Goose Birding World 10(1):11–15; an identification paper
Shields, Gerald F. (July 1990). "Analysis of mitochondrial DNA of Pacific Black Brant". The Auk. American Ornithologists' Union. 107 (3): 620–623.
Syroechkovski, E.E.; Zöckler, C.; Lappo, E. (December 1998). "Status of Brent Goose in northwestern Yakutia, East Siberia" (PDF). British Birds. British Birds Rarities Committee. 91 (12): 565–572. (this paper presented claims that Black Brant and Dark-bellied Brent Goose were interbreeding extensively in the Russian Arctic)
Sangster, George (2000). Taxonomic status of bernicla and nigricans Brent Goose British Birds 91(12):565–572 (a critical re-evaluation of the claims made in the above paper by Syroechkovski et al.)
Ebels, E.B. (1997). "Identification of brent geese: a new feature" (PDF). Dutch Birding. Dutch Birding Association. 19 (5): 232–236 (highlights the differences in belly colour between the various forms).

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