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Eugenes fulgens

Eugenes fulgens

Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Superphylum: Deuterostomia
Phylum: Chordata
Subphylum: Vertebrata
Infraphylum: Gnathostomata
Megaclassis: Osteichthyes
Cladus: Sarcopterygii
Cladus: Rhipidistia
Cladus: Tetrapodomorpha
Cladus: Eotetrapodiformes
Cladus: Elpistostegalia
Superclassis: Tetrapoda
Cladus: Reptiliomorpha
Cladus: Amniota
Classis: Reptilia
Cladus: Eureptilia
Cladus: Romeriida
Subclassis: Diapsida
Cladus: Sauria
Infraclassis: Archosauromorpha
Cladus: Crurotarsi
Divisio: Archosauria
Cladus: Avemetatarsalia
Cladus: Ornithodira
Subtaxon: Dinosauromorpha
Cladus: Dinosauriformes
Cladus: Dracohors
Cladus: Dinosauria
Ordo: Saurischia
Cladus: Eusaurischia
Subordo: Theropoda
Cladus: Neotheropoda
Cladus: Averostra
Cladus: Tetanurae
Cladus: Avetheropoda
Cladus: Coelurosauria
Cladus: Tyrannoraptora
Cladus: Maniraptoromorpha
Cladus: Maniraptoriformes
Cladus: Maniraptora
Cladus: Pennaraptora
Cladus: Paraves
Cladus: Eumaniraptora
Cladus: Avialae
Infraclassis: Aves
Cladus: Euavialae
Cladus: Avebrevicauda
Cladus: Pygostylia
Cladus: Ornithothoraces
Cladus: Ornithuromorpha
Cladus: Carinatae
Parvclassis: Neornithes
Cohors: Neognathae
Cladus: Neoaves
Superordo: Caprimulgimorphae
Ordo: Apodiformes

Familia: Trochilidae
Subfamilia: Trochilinae
Genus: Eugenes
Species: E. fulgens

Eugenes fulgens, (Swainson, 1827)

Philos.Mag.n.s. 1 p.441

Vernacular names
English: Magnificent Hummingbird
français: Colibri de Rivoli
polski: Ametyścik

Rivoli's hummingbird (Eugenes fulgens) is a species of hummingbird in the "mountain gems", tribe Lampornithini in subfamily Trochilinae. It is found in El Salvador, Guatemala, Honduras, Mexico, Nicaragua, and the United States.[3][4]

Taxonomy and systematics

Eugenes fulgens was originally described as the magnificent hummingbird, and by the late 1800s was treated as having two subspecies. Beginning in 2017 the North American Classification Committee of the American Ornithological Society (NACC), the International Ornithological Committee (IOC), and the Clements taxonomy split them into the current Rivoli's hummingbird and the Talamanca hummingbird (E. spectabilis). However, as of 2020, BirdLife International's Handbook of the Birds of the World (HBW) retains the single species "magnificent" hummingbird treatment.[5][3][6][4]

Rivoli's hummingbird is monotypic.[3]

Rivoli's hummingbird is about 11 to 14 cm (4.3 to 5.5 in) long and weighs about 6 to 10 g (0.21 to 0.35 oz), with males typically a little larger than females. It has a wingspan of 18 centimetres (7.1 in). Of the hummingbirds found in the United States, Rivoli's hummingbird is one of the two largest species, rivaled in size only by the blue-throated hummingbird. The black bill is long and straight to slightly decurved. Both sexes look very dark unless the sun catches the iridescence of the plumage and the brilliant colors flash in the sunlight. During much of the year the species is in the "definitive basic" breeding plumage described below. It has a protracted molt between that plumage and the duller non-breeding appearance. There are some minor clinal variations in bill length and the color of the vent area.[7][8]

The adult male is green-bronze dorsally, becoming more bronzed on the black-tipped tail. The crown is violet, the throat gorget bright blue-green, and the rest of the head black apart from a white spot behind the eye. The chest is green-bronze and the belly grayish. The female is bronze-green dorsally and has a dull gray ventral coloring. There is a white stripe behind her eye. Immature birds are like the female, but darker and browner.[7]
Distribution and habitat

Rivoli's hummingbird breeds in mountains from the southwestern United States to Honduras and Nicaragua. It inhabits the interior, edges, and clearings of pine-oak forest and cloudforest from about 1,500 m (4,900 ft) up to the timberline as well as more open secondary forest and some grasslands. It regularly comes to bird feeders.[7]

Rivoli's hummingbird is migratory in part of its range. It is a year-round resident from north-central Mexico south to northern Nicaragua. It nests but does not overwinter in southeastern Arizona, possibly in southwestern New Mexico, and the Mexican states of Chihuahua, Sonora, and possibly others. There are summer records in the Chisos Mountains of Texas but breeding has not been confirmed there.[7]
Interspecific interaction

Throughout its range Rivoli's hummingbird is subordinate to the larger blue-throated hummingbird. Rivoli's coexists with many other hummingbird species including black-chinned (Archilocus alexandri) and broad-tailed (Selasphorus platycercus). In Mexico they tend to dominate smaller hummingbirds such as amethyst-throated (Lampornis amethystinus), berylline (Saucerottia beryllina), and bumblebee (Atthis heloisa).[7]

Rivoli's hummingbird feeds on nectar from a wide variety of flowering plants whose composition varies by season, elevation, and latitude. In parts of Mexico at least, males defend feeding territories. It also feeds on small insects, which apparently make a larger part of its diet than the diets of other North American hummingbirds. It captures them in mid-air or by gleaning from vegetation while hovering.[7]

The Rivoli's hummingbird breeding season spans from May to July in the U.S and is possibly year-round in El Salvador. It has not been defined elsewhere. It builds a open cup nest of soft feathers and moss bound with spider silk and covered with lichen. The cup's interior is about 1.9 to 3.2 cm (0.75 to 1.3 in) deep and 2.9 to 3.8 cm (1.1 to 1.5 in) wide. They are typically placed on a horizontal branch or in a fork, often in maples and sycamores, and often over streams. They are usually at least 6 m (20 ft) above the ground. The female alone incubates the clutch of two white eggs; the period is not known but is probably 15 to 19 days like that of many other hummingbirds. The time to fledging appears to be about 25 days.[7]

Dickcissel male perched on a metal pole singing, with neck stretched and beak open.

Songs and calls
Listen to Rivoli's hummingbird on xeno-canto

What might be Rivoli's hummingbird song is "a loud or high-pitched, sharp tchik or tcheep." Another possible song is "a short chatter of notes that rise and fall". Males give single or a string of chip notes both while perched and in flight. During aggressive actions or alarm, the species gives a very rapid sequence of chip notes.[7]

The IUCN follows HBW taxonomy and so treats Rivoli's and Talamanca hummingbirds as a single species assessed as being of Least Concern. As a whole it has a large range and an apparently stable population, though the population size is not known.[1] Forest fires are a potential threat in the U.S. because Rivoli's is found only in higher isolated mountain ranges. "Habitat destruction in southern Mexico and Central America may have a longer-lasting impact on populations".[7]

BirdLife International (2021). "Magnificent Hummingbird Eugenes fulgens". IUCN Red List of Threatened Species. 2016: e.T22687746A168972435. doi:10.2305/IUCN.UK.2021-3.RLTS.T22687746A168972435.en. Retrieved 11 May 2022.
"Appendices | CITES". Retrieved 2022-01-14.
Gill, F.; Donsker, D.; Rasmussen, P. (July 2021). "IOC World Bird List (v 12.1)". doi:10.14344/IOC.ML.11.2. Retrieved 15 January 2022.
HBW and BirdLife International (2020) Handbook of the Birds of the World and BirdLife International digital checklist of the birds of the world Version 5. Available at: [.xls zipped 1 MB] retrieved 27 May 2021
"Check-list of North and Middle American Birds". American Ornithological Society. June 29, 2021. Retrieved August 9, 2021.
Clements, J. F., T. S. Schulenberg, M. J. Iliff, S. M. Billerman, T. A. Fredericks, J. A. Gerbracht, D. Lepage, B. L. Sullivan, and C. L. Wood. 2021. The eBird/Clements checklist of Birds of the World: v2021. Downloaded from Retrieved August 25, 2021
Powers, D. R., R. Partida-Lara, and P. L. Enríquez (2020). Rivoli's Hummingbird (Eugenes fulgens), version 1.0. In Birds of the World (S. M. Billerman, Editor). Cornell Lab of Ornithology, Ithaca, NY, USA. retrieved 11 May 2022 "Colibri de Rivoli - Eugenes fulgens - Rivoli's Hummingbird". Retrieved 2020-09-30.

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