Ortalis guttata

Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Superphylum: Deuterostomia
Phylum: Chordata
Subphylum: Vertebrata
Infraphylum: Gnathostomata
Megaclassis: Osteichthyes
Cladus: Sarcopterygii
Cladus: Rhipidistia
Cladus: Tetrapodomorpha
Cladus: Eotetrapodiformes
Cladus: Elpistostegalia
Superclassis: Tetrapoda
Cladus: Reptiliomorpha
Cladus: Amniota
Classis: Reptilia
Cladus: Eureptilia
Cladus: Romeriida
Subclassis: Diapsida
Cladus: Sauria
Infraclassis: Archosauromorpha
Cladus: Crurotarsi
Divisio: Archosauria
Cladus: Avemetatarsalia
Cladus: Ornithodira
Subtaxon: Dinosauromorpha
Cladus: Dinosauriformes
Cladus: Dracohors
Cladus: Dinosauria
Ordo: Saurischia
Cladus: Eusaurischia
Subordo: Theropoda
Cladus: Neotheropoda
Cladus: Averostra
Cladus: Tetanurae
Cladus: Avetheropoda
Cladus: Coelurosauria
Cladus: Tyrannoraptora
Cladus: Maniraptoromorpha
Cladus: Maniraptoriformes
Cladus: Maniraptora
Cladus: Pennaraptora
Cladus: Paraves
Cladus: Eumaniraptora
Cladus: Avialae
Infraclassis: Aves
Cladus: Avebrevicauda
Cladus: Pygostylia
Cladus: Ornithothoraces
Cladus: Ornithuromorpha
Cladus: Carinatae
Parvclassis: Neornithes
Cohors: Neognathae
Cladus: Pangalloanserae
Cladus: Galloanseres
Ordo: Galliformes
Subordo: Craci

Familia: Cracidae
Genus: Ortalis
Species: Ortalis guttata
Subspecies: O. g. guttata – O. g. remota – O. g. subaffinis
Name

Ortalis guttata (Spix, 1825)

Type locality: Río Solimões, Brazil.

Synonyms

Penelope guttata (protonym)

References
Primary references

Spix, J.B. von 1825. Avium species novae, quas in itinere per Brasiliam annis MDCCCXVII-MDCCCXX jussu et auspiciis Maximiliani Josephi I. Bavariae regis. Suscepto. Collegit et descripsit Dr. J. B. de Spix. Tomus 2. 85 pp. + 115 tt. Illustrations: Matthias Schmidt. Ed. Typis Franc. Seraph. Hübschmanni. Monachii. DOI: 10.5962/bhl.title.63182 Original description p. 55 BHL Reference page. pl. 73 BHL

Additional references

Silveira, L.F., Tomotani, B.M., Cestari, C., Straube, F.C. & Piacentini, V.Q. 2017. Ortalis remota: a forgotten and critically endangered species of chachalaca (Galliformes: Cracidae) from Eastern Brazil. Zootaxa 4306(4): 524–536. DOI: 10.11646/zootaxa.4306.4.4 Reference page.

Vernacular names
English: Speckled Chachalaca
español: Chachalaca moteada
português: aracuã-pintado
svenska: Spräcklig chachalaca

The speckled chachalaca (Ortalis guttata) is a species of bird in the family Cracidae, the chachalacas, guans, and curassows. It is found in Bolivia, Brazil, Colombia, Ecuador, and Peru.[2][3]
Taxonomy and systematics

The taxonomic history of the speckled chachalaca is complex. It was at one time considered conspecific with what was then called the variable chachalaca (Ortalis motmot) and buff-browed chachalaca (O. superciliaris). They were split and the variable chachalaca has since been split further.[4] (Confusingly, the International Ornithological Committee (IOC) then renamed O. motmot little chachalaca but the American Ornithological Society (AOS) and the Clements taxonomy retained the name "variable" for it.) Later still, three more species were split from speckled chachalaca, the East Brazilian (O. araucuan), scaled (O. squamata), and Colombian (O. columbiana) chachalacas.[2][5][6]

The speckled chachalaca as now understood has three subspecies, the nominate O. g. guttata, O. g. subaffinis, and O. g. remota. However, some authors treat the last as a subspecies of scaled chachalaca and others suggest it deserves to be an entirely separate species.[2][4]
Description

The speckled chachalaca is 49 to 55 cm (1.6 to 1.8 ft) long. The nominate subspecies is mostly brown above with a darker and grayer head and neck. The mantle has a chestnut wash; the tail has a bronze gloss and the outer feathers are rufous. The throat and breast are dark brown with the eponymous white speckles. The lower breast and belly are light grayish brown and the vent bright rufous. The eye is dark brown to dark reddish brown and is surrounded by bare slate blue skin.[4]

O. g. subaffinis is lighter but browner overall than the nominate, does not have the chestnut wash on the mantle, and the marks on the throat and breast are more scalloped than speckled. It is also somewhat larger. O. g. remota's description is from a single specimen. It differs from the nominate: Its crown is rufescent rather than gray, it is paler and more olive above, the rump is chestnut, and the "speckles" are larger and less well defined.[4]
Distribution and habitat

The nominate subspecies of speckled chachalaca is found in the Amazonia of eastern Colombia, Ecuador, and Peru; northern Bolivia; and western Brazil. O. g. subaffinis is found in the Amazonia of eastern and northeastern Bolivia and adjoining southwestern Brazil. O. g. remota is, as its specific epithet implies, remote from the other two subspecies; it is found in southeastern Brazil's Mato Grosso do Sul and São Paulo states.[4]

The speckled chachalaca inhabits a variety of tropical and subtropical landscapes. The two Amazonian subspecies is typically found in low or open forest, thickets, "forest islands" in savanna, várzea, and riparian and floodplain forest. They shun the interior of terra firme forest. They reach their highest elevations at the western edge of their range, usually up to 2,000 m (6,600 ft) in Bolivia, 1,100 m (3,600 ft) in Ecuador, and 1,700 m (5,600 ft) in Peru though locally higher in all three countries. O. g. remota is found only in secondary woodland and the edges of gallery forest in its small zone.[4]
Behavior
Feeding

The speckled chachalaca mostly forages in small flocks of up to seven birds, primarily in the subcanopy and understory and only rarely on the ground. Its diet has not been extensively studied but is known to include fruits and other plant material.[4]
Breeding

The speckled chachalaca's breeding phenology is poorly known. Its nesting season appears to vary across its range but in Peru encompasses at least November to March. One nest was made entirely of leaves on a bed of dry ferns. The clutch size is three to four eggs.[4]

Dickcissel male perched on a metal pole singing, with neck stretched and beak open.

Songs and calls
Listen to speckled chachalaca on xeno-canto
Vocalization

The song of the nominate speckled chachalaca is a "rhythmic five-syllable phrase rendered ha-ga-GAA-gogok, repeated rapidly." Those of the other subspecies are similar.[4]
Status

The IUCN has assessed the speckled chachalaca as being of Least Concern.[1] The two Amazonian subspecies are generally common throughout their range and occur in protected areas in every country. They seem to tolerate human activity fairly well, and hunting pressure is less than that suffered by other chachalacas. O. g. remota, however, is very rare; much of its habitat was flooded by hydroelectric reservoirs and Brazilian authorities consider it Critically Endangered.[4]
References

BirdLife International (2019). "Speckled Chachalaca Ortalis guttata". IUCN Red List of Threatened Species. 2019. Retrieved 29 September 2021.
Gill, F.; Donsker, D.; Rasmussen, P. (July 2021). "IOC World Bird List (v 11.2)". Retrieved July 14, 2021.
Remsen, J. V., Jr., J. I. Areta, E. Bonaccorso, S. Claramunt, A. Jaramillo, D. F. Lane, J. F. Pacheco, M. B. Robbins, F. G. Stiles, and K. J. Zimmer. Version 24 August 2021. Species Lists of Birds for South American Countries and Territories. https://www.museum.lsu.edu/~Remsen/SACCCountryLists.htm retrieved August 24, 2021
Kirwan, G. M., J. del Hoyo, N. Collar, and D. A. Christie (2020). Speckled Chachalaca (Ortalis guttata), version 1.0. In Birds of the World (S. M. Billerman, B. K. Keeney, P. G. Rodewald, and T. S. Schulenberg, Editors). Cornell Lab of Ornithology, Ithaca, NY, USA. https://doi.org/10.2173/bow.specha3.01 retrieved September 29, 2021
Remsen, J. V., Jr., J. I. Areta, E. Bonaccorso, S. Claramunt, A. Jaramillo, D. F. Lane, J. F. Pacheco, M. B. Robbins, F. G. Stiles, and K. J. Zimmer. Version 24 August 2021. A classification of the bird species of South America. American Ornithological Society. https://www.museum.lsu.edu/~Remsen/SACCBaseline.htm retrieved August 24, 2021
Clements, J. F., T. S. Schulenberg, M. J. Iliff, S. M. Billerman, T. A. Fredericks, J. A. Gerbracht, D. Lepage, B. L. Sullivan, and C. L. Wood. 2021. The eBird/Clements checklist of Birds of the World: v2021. Downloaded from https://www.birds.cornell.edu/clementschecklist/download/ Retrieved August 25, 2021