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Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Superphylum: Deuterostomia
Phylum: Chordata
Subphylum: Vertebrata
Infraphylum: Gnathostomata
Megaclassis: Osteichthyes
Cladus: Sarcopterygii
Cladus: Rhipidistia
Cladus: Tetrapodomorpha
Cladus: Eotetrapodiformes
Cladus: Elpistostegalia
Superclassis: Tetrapoda
Cladus: Reptiliomorpha
Cladus: Amniota
Cladus: Synapsida
Cladus: Eupelycosauria
Cladus: Sphenacodontia
Cladus: Sphenacodontoidea
Cladus: Therapsida
Cladus: Theriodontia
Subordo: Cynodontia
Infraordo: Eucynodontia
Cladus: Probainognathia
Cladus: Prozostrodontia
Cladus: Mammaliaformes
Classis: Mammalia
Subclassis: Trechnotheria
Infraclassis: Zatheria
Supercohors: Theria
Cohors: Eutheria
Infraclassis: Placentalia
Cladus: Boreoeutheria
Superordo: Euarchontoglires
Ordo: Rodentia
Subordo: Myomorpha
Superfamilia: Muroidea

Familia: Cricetidae
Subfamilia: Sigmodontinae
Tribus: Akodontini
Genus: Scapteromys
Species (3 + 1†): S. aquaticus – †S. hershkovitzi – S. meridionalis – S. tumidus
Name

Scapteromys Waterhouse, 1837: 20

Type species: Mus tumidus Waterhouse, 1837, by original designation.
References
Primary references

Waterhouse, G.R. 1837. Characters of new species of the genus Mus, from the collection of Mr. Darwin. Proceedings of the Zoological Society of London 1837(part V): 15–21, 27–32.

Additional references

Hershkovitz, P. 1966. South American swamp and fossorial rats of the Scapteromyine Group (Cricetinae, Muridae) with comments on the glans penis in murid taxonomy. Zeitschrift für Säugetierkunde 31: 81–149.
D′Elía, G. & Pardiñas, U.F.J. 2004. Systematics of Argentinean, Paraguayan, and Uruguayan swamp rats of the genus Scapteromys (Rodentia, Cricetidae, Sigmodontinae). Journal of Mammalogy 85(5): 897–910. DOI: 10.1644/BRB-201
Bonvicino, C.R., Fernandes, F.A., Viana, M.C., Teixeira, B.R. & D’Andrea, P.S. 2013. Scapteromys aquaticus (Rodentia: Sigmodontinae) in Brazil with comments on karyotype and phylogenetics relationships. Zoologia 30(2): 242–247. DOI: 0.1590/S1984-46702013000200016
Pardiñas, U.F.J. 2013. New Data on Scapteromys hershkovitzi Reig, 1994 (Rodentia: Cricetidae) from the Pliocene of Argentina. Ameghiniana 50(6): 593–597. DOI: 10.5710/AMGH.27.09.2013.642 Full article (PDF)
D’Elía, G. & Pardiñas, U.F.J. 2015. Genus Scapteromys Waterhouse, 1837. Pp. 269–274 in Patton, J.L., Pardiñas, U.F.J. & D’Elía, G. (eds.), Mammals of South America. Volume 2. Rodents. The University of Chicago Press: Chicago, Illinois. ISBN 978-0-226-16957-6. Reference page.

Links

Scapteromys in Mammal Species of the World.
Wilson, Don E. & Reeder, DeeAnn M. (Editors) 2005. Mammal Species of the World – A Taxonomic and Geographic Reference. Third edition. ISBN 0-8018-8221-4.

Vernacular names
English: Swamp Rat

Scapteromys is a genus of South American rodents in the tribe Akodontini of family Cricetidae. Three species are known, found in northern Argentina, southern Brazil, Paraguay and Uruguay.[1][2] They are as follows:

Argentine swamp rat (Scapteromys aquaticus)
Plateau swamp rat (Scapteromys meridionalis)
Waterhouse's swamp rat (Scapteromys tumidus)

Species are semiaquatic, living in and near marshes and other bodies of water. They reach a body length of 15 to 20 cm and a tail length of 13–17 cm, and weigh 110-200 g. Fur color is dark gray on top and light gray on the underside. They are primarily crepuscular and nocturnal. Their diet consists mainly of insects; they also consume other invertebrates and plant material.

The three species differ in karyotype, with aquaticus having 2n = 32, tumidus 2n = 24[1] and meridionalis 2n = 34/36.[2]
References

Musser, G.G.; Carleton, M.D. (2005). "Superfamily Muroidea". In Wilson, D.E.; Reeder, D.M (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference (3rd ed.). Johns Hopkins University Press. p. 1172. ISBN 978-0-8018-8221-0. OCLC 62265494.

Quintela, F.M.; Goncalves, G.L.; Althoff, S.L.; Sbalqueiro, I.J.; Oliveira, L.F.B.; de Freitas, T.R.O. (2014). "A new species of swamp rat of the genus Scapteromys Waterhouse, 1837 (Rodentia: Sigmodontinae) endemic to Araucaria angustifolia forest in southern Brazil". Zootaxa. 3811 (2): 207–225. doi:10.11646/zootaxa.3811.2.3. PMID 24943159.

Nowak, Ronald M. (1999). Walker's Mammals of the World. Vol. 2 (6th ed.). Johns Hopkins University Press. pp. 1398–1399. ISBN 978-0-8018-5789-8. OCLC 39045218.

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