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Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Spiralia
Cladus: Lophotrochozoa
Phylum: Mollusca
Classis: Gastropoda
Subclassis: Caenogastropoda
Ordo: Littorinimorpha
Superfamilia: Stromboidea

Familia: Aporrhaidae
Genera (3 + 73†): Aporrhais – Arrhoges – Cuphosolenus – †Alarimella – †Anchura – †Antarctohoges – †Araeodactylus – †Auriala – †Austroaporrhais – †Bicorempterus – †Biculteriala – †Bizarrus – †Ceratosiphon – †Checchiaia – †Cultrigera – †Cuphotifer – †Cyclomolops – †Diarthema – †Dicroloma – †Diempterus – †Digitolabrum – †Dimorphosoma – †Disoketa – †Drepanocheilus – †Ectinochilus – †Goniocheila – †Graciliala – †Gymnarus – †Harpagodes – †Harpodactylus – †Harpospira – †Helicaulax – †Hemichenopus – †Kangilioptera – †Kaunhowenia – †Latiala – †Lispodesthes – †Maussenetia – †Mexopus – †Monocuphus – †Myurusina – †Perissoptera – †Peruchilus – †Perustrombus – †Phyllocheilus – †Pietteia – †Platyoptera – †Pletzachia – †Pseudanchura – †Pterocerella – †Pugioptera – †Pugnellus – †Pyktes – †Quadrinervus – †Rhombovomeria – †Rimetella – †Spinigera – †Spiniloma – †Strombites – †Strombopugnellus – †Struthiochenopus – †Struthioptera – †Teneposita – †Tephlon – †Tessarolax – †Tibiaporrhais – †Toarctocera – †Torgnellus – †Tridactylus – †Trilemma – †Tulochilus – †Tundora – †Ueckeritzella – †Undoriptera – †Volgacheilus – †Wateletia

[Sources: GBIF for all above genera; BioLib lists 47 of the above genera; WoRMS only shows 15 of the above genera; all three sources are as of July, 2019; further investigation is needed to determine the true genera situation]
Name

Aporrhaidae Gray, 1850
Synonyms

Aporrhaididae (misspelling)
Aporrhaiidae (Spelling variation)
Arrhoginae Popenoe, 1983
Chenopidae Deshayes, 1865
Dicrolomatidae Korotkov, 1992 †
Perissopterinae Korotkov, 1992 †
Pugnellidae Kiel & Bandel, 1999 † (original rank)
Tundorinae Bandel, 2007 †

References

Kronenberg, G.C. 1991: The Recent species of the family Aporrhaidae. Vita marina, 41(2): 73–84. [not seen]

Links

Global Biodiversity Information Facility. 2019. GBIF Backbone Taxonomy. Checklist dataset. Taxon: Aporrhaidae.
Zicha, Ondřej et al. Aporrhaidae – Taxon details on Biological Library (BioLib).
Aporrhaidae in the World Register of Marine Species

Aporrhaidae is a family of sea snails commonly called the "pelican's foot snails." The taxonomy of the Gastropoda by Bouchet & Rocroi, 2005[3] categorizes Aporrhaidae as marine gastropod mollusks in the clade Littorinimorpha.

Aporrhaids are commercially important, especially in traditional fisheries.[4]

Description

Aporrhaids have a large lip with finger-like extensions, and a small operculum. They do not have movable eyes. Instead, their eyes are fixed at the base of each tentacle.[5]

The mollusk has one narrow foot, which renders its motion interrupted as the foot must raise the shell a tiny bit in each single movement, simultaneously pushing it forward. The motion must be repeated again and again for the mollusk to travel.[6]
Habitat

Aporrhaidae live on muddy and sandy bottoms, sometimes in very large populations.
Subfamilies

Subfamilies in the family Aporrhaidae include:

Aporrhainae Gray, 1850 - synonym: Chenopidae Deshayes, 1865
Arrhoginae Popenoe, 1983 - synonyms: Alariidae Koken, 1889 (inv.); Dicrolomatidae Korotkov, 1992
† Harpagonidae Pchelintsev, 1963
† Perissopterinae Korotkov, 1992 - synonym: Struthiopterinae Zinsmeister & Griffin, 1995
† Spinigerinae Korotkov, 1992 (inv.)

Aporrhais from the Pliocene of Cyprus.
Genera

There are only two Recent genera:[7][8]

Aporrhais da Costa, 1778
Arrhoges Gabb, 1868
† Dicroloma Gabb, 1868
† Drepanocheilus Meek, 1864
† Hemichenopus Steinmann & Wilckens, 1908
† Struthioptera Finlay & Marwick, 1937

Fossil genera within the family Aporrhaidae include:

Alarimella Saul, 1998
Anchura Conrad, 1860
Antarctohoges Stilwell & Zinsmeister, 1992
Araeodactylus Harris & Burrows, 1891
Auriala Hacobjan, 1976
Austroaporrhais Zinsmeister & Griffin, 1995
Bicorempterus Gründel, 2001
Ceratosiphon Gill, 1870
Cultrigera Böhm, 1885
Cuphosolenus Piette, 1876
Cuphotifer Piette, 1876
Cyclomolops Gabb, 1868
Diarthema Piette, 1864
Digitolabrum Cossmann, 1904
Dimorphosoma Gardner, 1875
Graciliala Sohl, 1960
Goniocheila Gabb, 1868
Helicaulax Gabb, 1868
Kangilioptera Rosenkrantz, 1970
Kaunhowenia Abdel Gawad, 1986
Latiala Sohl, 1960
Lispodesthes White, 1875
Maussenetia Cossmann, 1904
Mexopus Kiel & Perrilliat, 2001
Monocuphus Piette, 1876
Perissopter Tate, 1865
Peruchilus Olsson, 1931
Pietteia Cossmann, 1904
Platyoptera Conrad, 1855
Pseudanchura Kollmann, 2005
Pterocerella Meek, 1864
Pugioptera Pchelincev, 1953
Strombopugnellus Koch, 1911
Struthiochenopus Zinsmeister & Griffin, 1995
Teneposita Loch, 1989
Tessarolax Gabb, 1864
Tibiaporrhais Elder, 1990
Toarctocera Gründel, Nützel & Schulbert, 2009[9] - type species: Rostellaria subpunctata - Toarctocera subpunctata (von Munster in Goldfuss, 1826–1844)
Trilemma Blagovetshenskiy & Shumilkin, 2006
Tulochilus Finlay & Marwick, 1937
Tundora Stephenson, 1941
Ueckeritzella Gründel, 1998
Wateletia Cossmann, 1889

References

Gray J. E. 1850. Figures of molluscous animals selected from various authors. Etched for the use of students by M. E. Gray. Volume 4. Longman, Brown, Green & Longmans, London. iv+219 pp. Apporhaidae at page 66.
Kiel S. & Bandel K. (1999). "The Pugnellidae, a new stromboidean family from the Upper Cretaceous". Paläontologische Zeitschrift 73(1-2): 47-58, 6 Figs., Stuttgart. PDF[permanent dead link]
Bouchet P. & Rocroi J.-P. (Ed.); Frýda J., Hausdorf B., Ponder W., Valdes A. & Warén A. 2005. Classification and nomenclator of gastropod families. Malacologia: International Journal of Malacology, 47(1-2). ConchBooks: Hackenheim, Germany. ISBN 3-925919-72-4. ISSN 0076-2997. 397 pp. http://www.vliz.be/Vmdcdata/imis2/ref.php?refid=78278
"Shells aporrhaidae". Archived from the original on 2005-03-05. Retrieved 2009-03-07.
"Aporrhaidae (Pelicans Foot)". Archived from the original on 2019-02-22. Retrieved 2009-03-07.
"TrekNature | A Pelican's foot for Ahmet Photo".
Gofas, S. (2012). Aporrhaidae. Accessed through: World Register of Marine Species at http://www.marinespecies.org/aphia.php?p=taxdetails&id=22988 on 2012-09-27
Kronenberg, G.C. 1991: The Recent species of the family Aporrhaidae. Vita marina, 41(2): 73-84.

Gründel, J., Nützel A. & Schulbert C. (24 July) 2009. Toarctocera (Gastropoda, Aporrhaidae): a new genus from the Jurassic (Toarcian/Aalenian) of South Germany and the early evolutionary history of the family Aporrhaidae. Paläontologische Zeitschrift (online), PDF

Vaught, K.C. (1989). A classification of the living Mollusca. American Malacologists: Melbourne, FL (USA). ISBN 0-915826-22-4. XII, 195 pp.

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