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Classification System: APG IV

Superregnum: Eukaryota
Regnum: Plantae
Cladus: Angiosperms
Cladus: Eudicots
Cladus: Core eudicots
Cladus: Asterids
Cladus: Campanulids
Ordo: Asterales

Familia: Asteraceae
Subfamilia: Asteroideae
Tribus: Eupatorieae
Subtribus: Liatrinae
Genus: Carphephorus
Species: C. bellidifolius – C. carnosus – C. corymbosus – C. odoratissimus – C. paniculatus – C. pseudoliatris – C. tomentosus
Name

Carphephorus Cass., Bull. Sci. Soc. Philom. Paris 198. (1816)

Type species: Carphephorus pseudoliatris Cass., Bull. Sci. Soc. Philom. Paris 198. (1816)

Synonyms

Heterotypic
Litrisa Small (1924)
Type species: Litrisa carnosa Small (1924) vide Carphephorus carnosus (Small) C.W.James (1958)
Trilisa Cass.
Type species: Trilisa odoratissima (J.F. Gmel.) Cass. (1818) vide Carphephorus odoratissimus (J. Gmel.) Hebert (1968)

References

Cassini, A.H.G. de 1816. Bulletin des Sciences, par la Societe Philomatique : 198.
Hassler, M. 2018. Carphephorus. World Plants: Synonymic Checklists of the Vascular Plants of the World In: Roskovh, Y., Abucay, L., Orrell, T., Nicolson, D., Bailly, N., Kirk, P., Bourgoin, T., DeWalt, R.E., Decock, W., De Wever, A., Nieukerken, E. van, Zarucchi, J. & Penev, L., eds. 2018. Species 2000 & ITIS Catalogue of Life. Published on the internet. Accessed: 2018 January 05. Reference page.
International Plant Names Index. 2018. Carphephorus. Published online. Accessed: January 05 2018.
Tropicos.org 2018. Carphephorus. Missouri Botanical Garden. Published on the internet. Accessed: 2018 January 05.

Carphephorus is a genus of North American plants in the sunflower family.[1] They are native to the southeastern United States from Louisiana to Virginia.[2] Plants of this genus are known commonly as chaffheads.[3]

Description

These are perennial plants that grow from a caudex and fibrous root system. The stems are erect and unbranched, usually reaching 20 to 60 centimeters (8-24 inches) in height, and taller at times. The leaves are alternately arranged and point upward, sometimes pressed against the stem. The blades vary in shape and are hairy to hairless and generally glandular. The flower heads are borne in open inflorescences. Each head contains up to about 35 disc florets, usually lavender to dark magenta or pinkish purple, sometimes blue.[2] The fruit is a ribbed, rough-textured cypsela with a pappus of bristles.[2]
Classification

Some authors separate certain species into separate genera, Trilisa and Litrisa, on the basis of certain floral characters. The species are similar enough in other aspects that other authors maintain them in Carphephorus.[2] Molecular data may support the separation of at least some of the taxa.[4]

Carphephorus is in the tribe Eupatorieae of the aster family. Like other members of this tribe, the flower heads have disc florets and no ray florets. It is also in the subtribe Liatrinae along with, for example, Liatris and Garberia.[5]

Species and varieties[2][3][6][7]

Carphephorus bellidifolius – sandywoods chaffhead - Georgia South Carolina North Carolina Virginia
Carphephorus carnosus (syn. Trilisa carnosa) – pineland chaffhead - Florida
Carphephorus corymbosus – coastal plain chaffhead - Florida Georgia South Carolina
Carphephorus odoratissimus (syn. Trilisa odoratissima) – vanillaleaf - Louisiana Mississippi Alabama Florida Georgia South Carolina North Carolina
C. odoratissimus var. odoratissimus
C. odoratissimus var. subtropicanus (sometimes treated as a separate species, Carphephorus subtropicanus[8][9]). This species or variety is more southern than C. odoratissimus var. odoratissimus, lacks the characteristic coumarin odor of the latter, and has a different growth habit, being smaller with more of a rosette form. There is some overlap between the ranges of the two.[10]
Carphephorus pseudoliatris – bristleleaf chaffhead - Louisiana Mississippi Alabama Florida Georgia
Carphephorus tomentosus – woolly chaffhead - Georgia South Carolina North Carolina Virginia

formerly included[6]

Carphephorus paniculatus (syn of Trilisa paniculata ) – hairy chaffhead - Alabama Georgia Florida South Carolina North Carolina

Biochemistry

A number of species contain volatile oils, giving characteristic odors.[11][12]
References

Cassini, Alexandre Henri Gabriel de. 1816. Bulletin des Sciences, par la Societe Philomatique 1816: 198 in French
"Carphephorus Cassini". Flora of North America.
Carpephorus. Integrated Taxonomic Information System (ITIS).
Schilling, E. E.; P. B. Cox (2001). "Systematic analysis of Liatrinae (Asteraceae)". Botany 2001 Abstracts. Botanical Society of America.
"Garberia A.Gray". Flora of North America.
Flann, C (ed) 2009+ Global Compositae Checklist
Biota of North America Program 2013 county distribution maps
"Carphephorus odoratissimus var. subtropicanus". Atlas of Florida Vascular Plants.
Delaney K. R.; et al. (1999). "A new species of Carphephorus (Asteraceae; Eupatorieae) from peninsular Florida". Bot. Explor. (1).
Wunderlin, R. P.; Hansen, B. F. (2001). "Seven new combinations in the Florida flora". Novon. Missouri Botanical Garden Press. 11 (3): 366–369. doi:10.2307/3393048. JSTOR 3393048.
Karlsson, K. et al. (1972). Volatile constituents of Carphephorus corymbosus and Carphephorus paniculatus. Acta Chemica Scandinavica 26(10) 3839–48. ISSN 0001-5393
Karlsson, K.; et al. (1972). "Volatile constituents of Carphephorus odoratissimus (J.F. Gmel) Hebert". Acta Chemica Scandinavica. 26 (7): 2837–46. doi:10.3891/acta.chem.scand.26-2837. ISSN 0001-5393. PMID 4650326.

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