Hackelia

Life-forms

Classification System: APG IV

Superregnum: Eukaryota
Regnum: Plantae
Cladus: Angiosperms
Cladus: Eudicots
Cladus: Core eudicots
Cladus: Asterids
Cladus: Lamiids
Ordo: Boraginales

Familia: Boraginaceae
Subfamilia: Boraginoideae
Tribus: Eritrichieae
Genus: Hackelia
Species (57): H. amethystina – H. andicola – H. bella – H. besseyi – H. bhutanica – H. brachytuba – H. brevicula – H. californica – H. ciliata – H. cinerea – H. cronquistii – H. cusickii – H. davisii – H. deflexa – H. difformis – H. diffusa – H. floribunda – H. gracilenta – H. heliocapa – H. hendersonii – H. hintoniorum – H. hirsuta – H. hispida – H. ibapensis – H. latifolia – H. leonotis – H. macrophylla – H. meeboldii – H. mexicana – H. micrantha – H. mundula – H. murgabica – H. nervosa – H. obtusifolia – H. ophiobia – H. parviflora – H. patens – H. pinetorum – H. popovii – H. rattanii – H. revoluta – H. scaberrima – H. sessilifructa – H. setosa – H. sharsmithii – H. skutchii – H. stewartii – H. stricta – H. suaveolens – H. taylorii – H. uncinata – H. ursina – H. velutina – H. venusta – H. virginiana
Source(s) of checklist:

Govaerts, R. et al. 2020. Hackelia in Kew Science Plants of the World Online. The Board of Trustees of the Royal Botanic Gardens, Kew. Published online. Accessed: 2020 Sep 29. Reference page.

Name

Hackelia Opiz, 1839

Type species: Hackelia deflexa Opiz. LT designated by W. Punt, ING card 12780 (1960)

Synonyms

Austrocynoglossum Popov ex R.Mill.

References
Primary references

Opiz, P.M. in F.Berchtold, 1839. Oekonomisch-technische Flora Böhmens 2(2): 147

Additional references

Gentry, J.L. & Carr, R.L. 1976. A revision of the genus Hackelia (Boraginaceae) in North America, north of Mexico. Mem. New York Bot. Gard. 26(1): 121–227.
Johnston, I.M. 1923. Restoration of the genus Hackelia. Contr. Gray Herb. 68: 43–48.
Johnston, I.M. 1939. Studies in Boraginaceae. XIII. J. Arnold Arbor. 20: 575–402.
Bercht. & Opiz, Okon.-techn. Fl. Bohm. ii II. 146 (1838).

Links

Govaerts, R. et al. 2020. Hackelia in Kew Science Plants of the World Online. The Board of Trustees of the Royal Botanic Gardens, Kew. Published online. Accessed: 2020 Sep 29. Reference page.
International Plant Names Index. 2020. Hackelia. Published online. Accessed: Sep 29 2020. Reference page.
Tropicos.org 2020. Hackelia. Missouri Botanical Garden. Published online. Accessed: 29 Sep 2020. Reference page.

Catalogue of Life: 2024 Annual Checklist
USDA, ARS, Germplasm Resources Information Network. Hackelia in the Germplasm Resources Information Network (GRIN), U.S. Department of Agriculture Agricultural Research Service. Accessed: 08-Apr-12.

Vernacular names
English: stickseed
suomi: Korusirkunjyvät
中文: 假鹤虱属

Hackelia (stickseeds) is a genus of plants in the borage family, Boraginaceae. It includes 54 species found in North America, western South America, temperate Eurasia, and Australia.[1] 12 species are native to California.[2]

The genus was named after Josef Hackel, a Czech botanist. The common name, stickseed, refers to the tendency of the barbed nutlets to stick to animal fur.[3]
Species

54 species are currently accepted:[1]

Hackelia amethystina J.T.Howell - amethyst stickseed – California
Hackelia andicola (K.Krause) Brand – Peru
Hackelia bella (J.F.Macbr.) I.M.Johnst. - greater showy stickseed – northwestern California and southwestern Oregon
Hackelia besseyi (Rydb.) J.L. Gentry - Bessey's stickseed – Arizona, Colorado, New Mexico and Texas
Hackelia bhutanica R.R.Mill – eastern Nepal to northwestern Arunachal Pradesh
Hackelia brachytuba (Diels) I.M.Johnst. – Nepal to central China
Hackelia brevicula (Jeps.) J.L.Gentry - Poison Canyon stickseed – eastern California
Hackelia californica (A.Gray) I.M.Johnst. - California stickseed – northern California and west-central Oregon
Hackelia ciliata (Douglas ex Lehm.) I.M.Johnst. - British Columbia, Idaho, and Washington
Hackelia cinerea (Piper) I.M.Johnst. - gray stickseed – Washington, Idaho, and Montana
Hackelia cronquistii J.L.Gentry - Cronquist's stickseed – Oregon and Idaho
Hackelia cusickii (Piper) Brand - Cusick's stickseed – Oregon, northern California, and northwestern Nevada
Hackelia davisii Cronquist – Idaho
Hackelia deflexa (Wahlenb.) Opiz – northern Eurasia and northern North America
Hackelia difformis (Y.S.Lian & J.Q.Wang) Riedl – south-central China and Tibet
Hackelia diffusa (Douglas ex Lehm.) I.M.Johnst. - Spreading stickseed – British Columbia, Washington, Oregon
Hackelia floribunda (Lehm.) I.M.Johnst. - manyflower stickseed – western and central Canada and United States
Hackelia gracilenta I.M.Johnst. – Colorado
Hackelia heliocarpa (Brand) Brand – Mexico (Chihuahua)
Hackelia hendersonii (Piper) Brand – Washington, Oregon, and Montana
Hackelia hintoniorum (B.L.Turner) Sutorý – Mexico (Oaxaca)
Hackelia hirsuta (Wooton & Standl.) I.M.Johnst. – New Mexico
Hackelia hispida I.M.Johnst. - showy stickseed – Washington, Oregon, Idaho
Hackelia ibapensis L.M.Shultz & J.S.Shultz – Utah
Hackelia latifolia (R.Br.) Dimon & M.A.M.Renner – eastern Australia and Tasmania
Hackelia leonotis I.M.Johnst. – northeastern and central Mexico
Hackelia macrophylla (Brand) I.M.Johnst. – northern Pakistan and western Himalaya
Hackelia meeboldii Brand – western Himalaya
Hackelia mexicana (Schltdl. & Cham.) I.M.Johnst. – Mexico to Venezuela and Peru
Hackelia micrantha (Eastw.) J.L.Gentry - Jessica sticktight – western Canada and United States
Hackelia mundula (Jeps.) Ferris - pink stickseed – southwestern Oregon to central California
Hackelia murgabica Czukav. – Tajikistan
Hackelia nervosa (Kellogg) I.M.Johnst. - Sierra stickseed – California and northwestern Nevada
Hackelia obtusifolia R.R.Mill – Bhutan
Hackelia ophiobia R.L.Carr – southeastern Oregon, southwestern Idaho, and northern Nevada
Hackelia parviflora (K.Krause) Brand – Peru (Junín)
Hackelia patens (Nutt.) I.M.Johnst. - spotted stickseed – west-central United States
Hackelia pinetorum (Greene ex A.Gray) I.M.Johnst. – Arizona, New Mexico, Texas, and northern Mexico
Hackelia popovii Czukav. – Tajikistan
Hackelia rattanii (Brand) Brand – Mexico (Baja California)
Hackelia revoluta (Ruiz & Pav.) I.M.Johnst. – Colombia to northwestern Argentina
Hackelia sessilifructa (Y.S.Lian & J.Q.Wang) Ovczinnikova – Xinjiang
Hackelia setosa (Piper) I.M.Johnst. - bristly stickseed – northern California and southwestern Oregon
Hackelia sharsmithii I.M.Johnst. - Sharsmith's stickseed – central California and central Nevada
Hackelia skutchii I.M.Johnst. – Guatemala and Costa Rica
Hackelia stewartii I.M.Johnst. – northern Pakistan and western Himalaya
Hackelia stricta I.M.Johnst. – southwestern Mexico to Veracruz
Hackelia suaveolens (R.Br.) Dimon & M.A.M.Renner – eastern and southern Australia and Tasmania
Hackelia taylorii Harrod, Malmquist & R.L.Carr – Washington
Hackelia uncinata (Royle ex Benth.) C.E.C.Fisch. – northeastern Pakistan to Himalaya and southern China
Hackelia ursina (Greene ex A.Gray) I.M.Johnst. – southeastern Arizona, southwestern New Mexico, and northern Mexico (Sonora)
Hackelia velutina (Piper) I.M.Johnst. - velvet stickseed – north-central and central California and northwestern Nevada
Hackelia venusta (Piper) H.St.John - lesser showy stickseed – central Washington
Hackelia virginiana (L.) I.M.Johnst. - beggar's lice – central and eastern United States and eastern Canada

References

Hackelia Opiz. Plants of the World Online. Retrieved 17 June 2023.
Calflora Database: Hackelia species index in California. Calflora. Retrieved 17 June 2023.
Taylor, Ronald J. (1994) [1992]. Sagebrush Country: A Wildflower Sanctuary (rev. ed.). Missoula, MT: Mountain Press Pub. Co. p. 24. ISBN 0-87842-280-3. OCLC 25708726.