Primula veris

Classification System: APG IV

Superregnum: Eukaryota
Regnum: Plantae
Cladus: Angiosperms
Cladus: Eudicots
Cladus: Core eudicots
Cladus: Asterids
Ordo: Ericales

Familia: Primulaceae
Subfamilia: Primuloideae
Genus: Primula
Subgenus: P. subg. Primula
Section: P. sect. Primula
Species: Primula veris
Subspecies: P. v. subsp. canescens – P. v. subsp. columnae – P. v. subsp. macrocalyx – P. v. subsp. veris
Name

Primula veris L. (1753)
Synonyms

Primula veris Lehm., Monogr. Primul. 27. 1817, isonym.

Homonym

Primula veris Mill., Gard. Dict., ed. 8. n. 1. 1768, nom. illeg. non L. (1753), is synonymous to Primula vulgaris.
Hybrids

P. × media – P. × polyantha
References

Linnaeus, C. 1753. Species Plantarum 1: 142.
USDA, ARS, Germplasm Resources Information Network. Primula veris in the Germplasm Resources Information Network (GRIN), U.S. Department of Agriculture Agricultural Research Service. Accessed: 07-Oct-06.

Distribution

Native throughout most of temperate Europe and Asia although absent from more northerly areas including northern Scotland.
Vernacular names
azərbaycanca: Dərman novruzgülü
беларуская: Першацвет веснавы
български: Лечебна иглика
català: Cucut
kaszëbsczi: Kukówczé rãkawice
čeština: Prvosenka jarní
Cymraeg: Briallen Fair
dansk: Hulkravet Kodriver
Deutsch: Echte Schlüsselblume
English: Cowslip
Esperanto: Printempa primolo
español: Primicia de amistades
eesti: Harilik nurmenukk
فارسی: پریمولا وریس
suomi: Kevätesikko
français: Primevère officinale
Gaeilge: Bainne bó bleachtáin
hornjoserbsce: Wonjaty kropačk
magyar: Tavaszi kankalin
Ido: Primulo
italiano: Primavera
日本語: キバナノクリンザクラ
한국어: 황화구륜초
lietuvių: Pavasarinė raktažolė
latviešu: Gaiļbiksīte
Nedersaksies: Gullen sleutelbloeme
Nederlands: Gulden Sleutelbloem
norsk nynorsk: Marianøkleblom
norsk: Marianøkleblom
occitan: Cocut
polski: Pierwiosnek lekarski
русский: Первоцвет весенний
slovenčina: Prvosienka jarná
slovenščina: Pomladanski jeglič
svenska: Gullviva
українська: Первоцвіт весняний
中文: 黄花九轮草

Primula veris, the cowslip, common cowslip, or cowslip primrose (syn. Primula officinalis Hill), is a herbaceous perennial flowering plant in the primrose family Primulaceae. The species is native throughout most of temperate Europe and western Asia,[1] and although absent from more northerly areas including much of northwest Scotland, it reappears in northernmost Sutherland and Orkney and in Scandinavia.[2] This species frequently hybridizes with other Primulas such as the common primrose Primula vulgaris to form false oxlip (Primula × polyantha) which is often confused with true oxlip (Primula elatior), a much rarer plant.

Names

The common name cowslip may derive from the old English for cow dung, probably because the plant was often found growing amongst the manure in cow pastures.[3] An alternative derivation simply refers to slippery or boggy ground; again, a typical habitat for this plant.[4]

The species name veris (of spring, referring to the season) is the genitive case form of Latin "ver" (spring).[5] However, primrose P. vulgaris, flowers earlier, from December to May in the British Isles.[6]: 240 

Other historical common names include cuy lippe, herb peter, paigle or pagil,[7] peggle, key flower, key of heaven, fairy cups, petty mulleins, crewel, buckles, palsywort, and plumrocks.[8]
Description
Flowers
Primula veris – MHNT
Image of a trust of cowslips, gouache on vellum
Albrecht Dürer, Tuft of Cowslips, 1526, National Gallery of Art, Washington, D.C., NGA 74162

Primula veris is a variable evergreen or semi-evergreen perennial plant growing to 25 cm (10 in) tall and broad, with a rosette of leaves 5–15 cm long and 2–6 cm broad. The deep yellow flowers are produced in spring, in clusters of 10–30 blooms together on a single stem.[9] Each flower is 9–15 mm broad. Red- and orange-flowered plants occur rarely but can be locally widespread in areas where coloured primula hybrids bloom at the same time as the native cowslip, enabling cross-pollination.
Habitat and conservation
Primula veris in a meadow.

The cowslip is frequently found on more open ground than the primrose, including open fields, meadows, coastal dunes and clifftops. The plant suffered a decline due to changing agricultural practices throughout the 1970s and 1980s in Britain. It may therefore be rare locally, though where found it may be abundant. Additionally the seeds are now often included in wildflower seed mixes used to landscape motorway banks and similar civil engineering earthworks where the plants may be seen in dense stands. This practice has led to a revival in its fortunes.[10][11][12]
Cultivation

In cultivation this plant has gained the Royal Horticultural Society's Award of Garden Merit.[13][14]
Similar species
Red-flowered Primula veris plants

The cowslip may be confused with the closely related Primula elatior (oxlip) which has a similar general appearance and habitat, although the oxlip has larger, pale yellow flowers more like a primrose, and a corolla tube without folds.
Chemical constituents

The roots of Primula veris contain several glycosides of 5-methoxysalicylic methyl ester, such as primeverin[15] and primulaverin.[16] In the dried crude root, their phenolic aglycones are responsible for the typical odour reminiscent of methyl salicylate or anethole. The dried roots contain significant amounts of triterpene saponins, such as primula acid I/II, while in the flower these constituents are located in the sepals, and the dominating constituents are flavonoids.[17] Rare side effects of the saponins can be nausea or diarrhoea while some of the phenolic constituents are possibly responsible for allergic reactions.[18][19]

The subspecies macrocalyx, growing in Siberia, contains the phenolic compound riccardin C.[20]
Cuisine

Cowslip leaves have been traditionally used in Spanish cooking as a salad green. Uses in English cookery include using the flowers to flavor country wine[21] and vinegars; sugaring to be a sweet or eaten as part of a composed salad while the juice of the cowslip is used to prepare tansy for frying. The close cousin of the cowslip, the primrose P. vulgaris has often been confused with the cowslip and its uses in cuisine are similar with the addition of its flowers being used as a colouring agent in desserts.

English children's writer Alison Uttley in her story "The Country Child" (1931) of family life on an English farm from the perspective of a 9-year-old farmer's daughter Susan describes cowslips among the favourite flowers of her heroine and mentions her participation in preparing them for making cowslip wine, a locally important process. After its initial preparation, cowslip wine "would change to sparkling yellow wine" offered in "little fluted glasses" with a biscuit to important "morning visitors" of the farm: such as the curate coming for subscriptions, the local squire (landowner) and an occasional dealer (of their produce). This wine "was more precious than elderberry wine, which was the drink for cold weather, for snow and sleet".[21]
Myth and folklore

This herb was already mentioned by Pliny the Elder for its early blooming attributes. Species from the genus Primula along with other ritual plants played a significant role in the pharmacy and mythology of the Celtic druids, likely as an ingredient of magical potions to increase the absorption of other herbal constituents. In the Middle-Ages it was also known as St. Peter's herb or Petrella and was sought after by Florentine apothecaries. Hildegard von Bingen recommended the medicinal parts only for topical use but the leaves were also consumed as food. Other common names at the time were 'Herba paralysis', 'Verbascum', primrose or mullein leaves. It was frequently misidentified as or confused with similar species from the genus Primula.[22]
References

Anderberg, Arne. "Primula veris L." Naturhistoriska riksmuseet, Stockholm. Retrieved 1 March 2017.
Preston, Pearman & Dines (2002). New Atlas of the British Flora. Oxford University Press.
Anon. "Cowslip". Word-Origins. Word-Origins.com. Archived from the original on 18 July 2011. Retrieved 8 May 2010.
Shorter Oxford English dictionary, 6th ed. United Kingdom: Oxford University Press. 2007. p. 3804. ISBN 978-0199206872.
ver. Charlton T. Lewis and Charles Short. A Latin Dictionary on Perseus Project.
Clapham, A.R.; Tutin, T.G.; Warburg, E.F. (1981). Excursion flora of the British isles (3 ed.). Cambridge, U.K.: Cambridge University Press. ISBN 0521232902.
Notes to Beachy Head; with Other Poems: Charlotte Smith, 1807
Cowslip, from A Modern Herbal
RHS A-Z encyclopedia of garden plants. United Kingdom: Dorling Kindersley. 2008. p. 1136. ISBN 978-1405332965.
"Primula veris – Online Atlas of the British and Irish Flora". brc.ac.uk.
"Cowslip – Primula veris". naturespot.org.uk.
http://your.caerphilly.gov.uk/countryside/sites/your.caerphilly.gov.uk.countryside/files/pdf/actionplan/vol2-10-cowslip.pdf
"RHS Plant Selector – Primula veris". Retrieved 15 April 2020.
"AGM Plants – Ornamental" (PDF). Royal Horticultural Society. July 2017. p. 82. Retrieved 14 May 2018.
See CID 3038513 from PubChem for chemical structure
See Merck Index monograph for chemical details
Max Wichtl, ed. (2004). Herbal Drugs and Phytopharmaceuticals: A Handbook for Practice on a Scientific Basis. Medpharm Publishers. p. 473.
Howard, Michael (1987). Traditional Herbal Remedies. Century. pp. 128–129.
"Glykosidy" (in Czech). biotox.cz.
Yu. S. Kosenkova; M.P. Polovinka; N.I. Komarova; D.V. Korchagina; N. Yu. Kurochkina; V.A. Cheremushkina; N.F. Salakhutdinov (2009). "Seasonal Dynamics of Riccardin C Accumulation in Primula macrocalyx Bge" (PDF). Chemistry for Sustainable Development. 17: 507–511.[permanent dead link]
Uttley, Alison (1952). The Country Child. London: Faber and Faber. p. 156.
ES Burgess (Jan. 1902). History of Pre-Clusian Botany in Its Relation to Aster pgs 70, 72, 246, 324, 332, 334/5, 435, 440. Retrieved 19 December 2015.