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Classification System: APG IV

Superregnum: Eukaryota
Regnum: Plantae
Cladus: Angiosperms
Cladus: Monocots
Cladus: Commelinids
Ordo: Poales

Familia: Poaceae
Subfamilia: Puelioideae
Tribus: Guaduelleae - Puelieae
Genera: Guaduelleae - Puelieae

Puelioideae L.G. Clark, M. Kobay., S. Mathews, Spangler & E.A. Kellogg Syst. Bot. 25(2): 184–186. (2000)
Typus: Puelia Franch. Bull. Mens. Soc. Linn. Paris 1: 674. (1887)


Clark, L.G., Kobay, M., Mathews, Spangler, S. & Kellogg, E.A. 2000. Systematic Botany 25(2): 184–186.
Soreng, R.J., Peterson, P.M., Romaschenko, K., Davidse, G., Zuloaga, F.O., Judziewicz, E.J., Filgueiras, T.S., Davis, J.I. & Morrone, O. 2015. A worldwide phylogenetic classification of the Poaceae (Gramineae). Journal of Systematics and Evolution 53(2): 117–137. DOI: 10.1111/jse.12150 Paywall PDF. Reference page.
Simon, B.K., Clayton, W.D., Harman, K.T., Vorontsova, M., Brake, I., Healy, D. & Alfonso, Y. 2013. GrassWorld, Puelioideae. Published online. Accessed: 2013 Oct. 4. 2013. Puelioideae. Missouri Botanical Garden. Published online. Accessed: 3 Oct. 2013.
International Plant Names Index. 2013. Puelioideae. Published online. Accessed: 3 Oct. 2013.
USDA: GRIN Taxonomy for Plants[1] (23 March 2006)

The Puelioideae is a subfamily of the true grass family Poaceae with two genera, Guaduella and Puelia, each in its own tribe.[1] Its members grow in the understory of rainforests.

This subfamily is one of the earliest-diverging grass lineages, sister to the big BOP and PACMAD clades:[2]





BOP clade

PACMAD clade


Soreng, Robert J.; Peterson, Paul M.; Romaschenko, Konstantin; Davidse, Gerrit; Zuloaga, Fernando O.; Judziewicz, Emmet J.; Filgueiras, Tarciso S.; Davis, Jerrold I.; Morrone, Osvaldo (2015). "A worldwide phylogenetic classification of the Poaceae (Gramineae)". Journal of Systematics and Evolution. 53 (2): 117–137. doi:10.1111/jse.12150. ISSN 1674-4918. S2CID 84052108. open access
Grass Phylogeny Working Group II (2012). "New grass phylogeny resolves deep evolutionary relationships and discovers C4 origins". New Phytologist. 193 (2): 304–312. doi:10.1111/j.1469-8137.2011.03972.x. hdl:2262/73271. PMID 22115274. open access

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