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Classification System: APG IV

Superregnum: Eukaryota
Regnum: Plantae
Cladus: Angiosperms
Cladus: Eudicots
Cladus: Core eudicots
Cladus: Asterids
Ordo: Ericales

Familia: Primulaceae
Subfamilia: Theophrastoideae
Tribus: Samoleae
Genus: Samolus
Species: S. alyssoides – S. americanus – S. caulescens – S. cuneatus – S. ebracteatus – S. floribundus – S. latifolius – S. parviflorus – S. porosus – S. repens – S. spathulatus – S. subnudicaulis – S. vagans – S. valerandi

Samolus L. Sp. Pl. 1: 171 (1753).
Primary references

Linnaeus, C. 1753. Species Plantarum. Tomus I: 171. Reference page.

Additional references

Crusio W.E. 1982. Het geslacht Samolus L. Communications of the Dutch Waterplant Society 2: 13–25. (Dutch) Reference page.
Crusio W.E. 1984. Notes on the genus Samolus L. (Primulaceae). Communications of the Dutch Waterplant Society 6: 13–16. Reference page.

Vernacular names
Deutsch: Bunge
suomi: Suolapungat

Samolus (known as brookweed, or water pimpernel) is a widely distributed genus of about a dozen species of water-loving herbs. According to the APG III classification, this genus belongs to the family Primulaceae in the order Ericales. It was considered as closely related to a clade comprising the Theophrastaceae,[1] and was treated as part of that family or in its own monogeneric family, the Samolaceae. The APG III system does not recognize these families and instead includes all genera formerly belonging to Theophrastaceae in the family Primulaceae.

The last complete taxonomic treatment of this genus recognizes the following species:[2][3]

Samolus cinerascens
Samolus dichondrifolius
Samolus ebracteatus
Samolus junceus
Samolus porosus
Samolus pyrolifolius
Samolus repens
Samolus spathulatus
Samolus subnudicaulis
Samolus vagans
Samolus valerandi

Although some local floras recognize the North-American populations of S. valerandi as a separate species (S. parviflorus) or subspecies (S. valerandi ssp. parviflorus),[4] molecular and morphological data indicate that S. vagans and S. parviflorus should not be regarded as separate species but as part of a widespread "S. valerandi species complex.[5]

Caris PL, Smets EF (2004). "A floral ontogenetic study on the sister group relationship between the genus Samolus (Primulaceae) and the Theophrastaceae". American Journal of Botany. 91 (5): 627–643. doi:10.3732/ajb.91.5.627. PMID 21653418.
Crusio WE (20 February 1982). "Het geslacht Samolus L". Communications of the Dutch Waterplant Society (in Dutch). 2: 13–25.
Crusio WE (26 May 1984). "Notes on the genus Samolus L. (Primulaceae)". Communications of the Dutch Waterplant Society. 6: 13–16.
"Samolus". Flora of North America. Retrieved 2018-02-01.

Jones, K.; Anderberg, A. A.; Ronse De Craene, L. P.; Wanntorp, L. (23 June 2012). "Origin, diversification, and evolution of Samolus valerandi (Samolaceae, Ericales)". Plant Systematics and Evolution. 298 (8): 1523–1531. doi:10.1007/s00606-012-0655-z.


Wanntorp, Livia; Anderberg, Arne A. (2011). "Evolution And Diversification Of Brook Weeds (Samolus, Samolaceae, Ericales)". International Journal of Plant Sciences. 172 (2): 250–266. doi:10.1086/657647. JSTOR 10.1086/657647. S2CID 84493389.

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