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Classification System: APG IV

Superregnum: Eukaryota
Regnum: Plantae
Cladus: Angiosperms
Cladus: Eudicots
Cladus: Core eudicots
Cladus: Rosids
Cladus: Eurosids I
Ordoo: Rosales

Familia: Rosaceae
Subfamilia: Rosoideae
Tribus: Agrimonieae
Subtribus: Sanguisorbinae
Genera: AcaenaBencomiaCliffortiaMarcetellaMargyricarpusPolylepisPoteridiumPoteriumSanguisorbaSarcopoteriumTetraglochin


Sanguisorbinae Torr. & A.Gray, Fl. N. Amer. 1(3): 428 (1840).

Type genus: Sanguisorba L.

Note: The traditional tribal circumscription is to follow Sanguisorba s.l. with the addition of a small number of distinctive genera. However, Sanguisorba s.l. is paraphyletic with respect to Poteridium Spach, Ann. Sci. Nat., Bot., sér. 3, 5: 43 (1846) and Poterium L., Sp. Pl.: 994 (1753), which would both need to be resurrected (Potter et al., 2007 and Wang et al., 2020), if monophyly is required for this tribe.
Primary references

Torrey, J. & Gray, A. 1838–1840. A Flora of North America containing abridged Descriptions of all the known indigenous and naturalized Plants growing North of Mexico; arranged according to the Natural System, Vol. 1. 740 pp. Wiley & Putnam, New-York. BHL Reference page.

Additional references

Chen, X., Li, J., , Zhang, W., Liu, Y., Wu, P., Yang, X., Wang, L. & Zhou, S.-L. 2020. Molecular systematics of Rosoideae (Rosaceae). Plant Systematics and Evolution 306(1, 9): 1–12. DOI: 10.1007/s00606-020-01629-z Paywall ResearchGate Reference page.
Eriksson, T., Hibbs, M.S., Yoder, A.D., Delwiche, C.F., & Donoghue, M.J. 2003. The phylogeny of Rosoideae (Rosaceae) based on sequences of the internal transcribed spacers (ITS) of nuclear ribosomal DNA and the trnL/F region of chloroplast DNA. International Journal of Plant Sciences 164 (2): 197–211. DOI: 10.1086/346163 Full text PDF Reference page.
Potter, D., Eriksson, T., Evans, R.C., Oh, S., Smedmark, J.E.E., Morgan, D.R., Kerr, M., Robertson, K.R., Arsenault, M., Dickinson, T.A. & Campbell, C.S. 2007. Phylogeny and classification of Rosaceae. Plant Systematics and Evolution 266(1-2): 5–43. DOI: 10.1007/s00606-007-0539-9 PDF Reference page.
Wang, J.R., Wang, X., Su, N., Li, Q.J., Zhang, X.H., Ma, Y.P., Zhao, L., Ginefra Toni, J.F. & De Craene, L.R. 2020. Floral morphology and morphogenesis in Sanguisorba (Rosaceae): flower diversification despite petal reduction and spatial constraints. Botanical Journal of the Linnean Society 193(1): 47-63. DOI: 10.1093/botlinnean/boaa009 ResearchGate Reference page.


Stevens, P.F. 2001 onwards. Angiosperm Phylogeny Website. Version 14, July 2017 [and more or less continuously updated since]. Online. Reference page.

Vernacular names
العربية: مرقئينة

Sanguisorbinae is a subtribe of flowering plants in the rose family, Rosaceae. It is the sister to subtribe Agrimoniinae in tribe Sanguisorbeae.[1]

Eriksson, Torsten; Hibbs, Malin S.; Yoder, Anne D.; Delwiche, Charles F.; Donoghue, Michael J. (2003). "The Phylogeny of Rosoideae (Rosaceae) Based on Sequences of the Internal Transcribed Spacers (ITS) of Nuclear Ribosomal DNA and the TRNL/F Region of Chloroplast DNA". International Journal of Plant Sciences. 164 (2): 197–211 abstract. doi:10.1086/346163.

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