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Classification System: APG IV

Superregnum: Eukaryota
Regnum: Plantae
Cladus: Angiosperms
Cladus: Eudicots
Cladus: Core eudicots
Ordo: Caryophyllales

Familia: Caryophyllaceae
Tribus: Sperguleae
Genus: Spergularia
Species: S. aberrans – S. andina – S. angustata – S. arbuscula – S. atrosperma – S. azorica – S. bocconei – S. bungeana – S. canadensis – S. capillacea – S. collina – S. colombiana – S. confertiflora – S. congestifolia – S. cremnophila – S. denticulata – S. diandra – S. diandroides – S. doumerguei – S. echinosperma – S. embergeri – S. fasciculata – S. fimbriata – S. floribunda – S. hanoverensis – S. heldreichii – S. kurkae – S. lycia – S. macrorrhiza – S. macrotheca – S. manicata – S. masafuerana – S. media – S. melanocaulos – S. mexicana – S. microsperma – S. munbyana – S. nesophila – S. nicaeensis – S. pazensis – S. pitardiana – S. purpurea – S. pycnantha – S. pycnorrhiza – S. rubra – S. rupicola – S. salina – S. salontana – S. segetalis – S. sezer-zenginii – S. spruceana – S. stenocarpa – S. syvaschica – S. tangerina – S. tasmanica – S. tenuifolia

Spergularia (Pers.) J.Presl & C.Presl Fl. Cech. 94. (1819) nom. cons.

Type species: Spergularia rubra J.Presl & C.Presl Fl. Cech. 94. (1819)


Arenaria subg. Spergularia Pers., Syn. Pl. 1: 504. (1805)
Buda Adans., Fam. 2: 507, 528. (1763), nom. rej.
Type species: B. rubra (L.) Desv.
Tissa Adans., Fam. 2: 507, 611. (1763), nom. rej.
Type species: T. rubra (L.) Britton
Stipularia Haworth, Syn. Pl. Succ. 103. (1812), nom. illeg. non P.Beauv. (1810).
Type species: S. rubra Haw.
Lepigonum (Fr.) Wahlb., Fl. Gothob. 45. (1820)
Type species: L. rubrum Wahlb.
Melargyra Raf., Fl. Tell. 3: 81. 1836 publ. (1837)
Type species: M. rubra Raf.
Fasciculus Dulac, Fl. Hautes-Pyrénées 245. (1867)
Type species: F. ruber Dulac
Arenaria sect. Lepigonum Fr., Fl. Hall. 7-10: 159. (1819)
Alsine L. Sp. Pl. 1: 272. (1753)
Type species: A. media L.
Segetella Desv., J. Bot. Agric. 3: 255. (1816), nom. illeg. non Hedwig (1806).
Type species: S. vulgaris Desv.
Delia Dumort., Fl. Belg. 110. (1827)
Type species: D. segetalis Dumort.
Sanctambrosia Skottsb.apud Kuschel, Ark. Bot. ser. 2, 4: 418. (1962)

Note: This genus is not subsumed into Spergula.
Primary references

Presl, J.S. & Presl, C.B. 1819. Flora Čechica 94.

Additional references

Adams, L.G., West, J.G. & Cowley, K.J. 2008. Revision of Spergularia (Caryophyllaceae) in Australia. Australian Systematic Botany 21(4): 251–270. DOI: 10.1071/SB08015 Reference page.
Hernández-Ledesma, P., Berendsohn, W. G., Borsch, T., Mering, S. v., Akhani, H., Arias, S., Castañeda-Noa, I., Eggli, U., Eriksson, R., Flores-Olvera, H., Fuentes-Bazán, S., Kadereit, G., Klak, C., Korotkova, N., Nyffeler R., Ocampo G., Ochoterena, H., Oxelman, B., Rabeler, R. K., Sanchez, A., Schlumpberger, B. O. & Uotila, P. 2015. A taxonomic backbone for the global synthesis of species diversity in the angiosperm order Caryophyllales. Willdenowia 45(3): 281–383. DOI: 10.3372/wi.45.45301 Open access Reference page.
Kool, A. & Thulin, M. 2017. A giant spurrey on a tiny island: On the phylogenetic position of Sanctambrosia manicata (Caryophyllaceae) and the generic circumscriptions of Spergula, Spergularia and Rhodalsine. Taxon 66(3): 615-622. DOI: 10.12705/663.6 Reference page.
López González, G. 2010. Sobre el género Spergula L. [incl. Spergularia (Pers.) Pers. ex C. Presl & J Presl, nom. cons.] (Caryophyllaceae) y sus especies en la Península Ibérica e Islas Baleares. Lagascalia 30: 7–18 [1]. Proposal to include Spergularia (Pers.) Pers. ex C. Presl & J Presl into this genus is rejected.
Wilcox, M. 2009. Spergularia in Auckland. Auckland Botanical Society Journal 64: 73–74. Full article (PDF) Reference page.


Hassler, M. 2019. Spergularia. World Plants: Synonymic Checklists of the Vascular Plants of the World In: Roskovh, Y., Abucay, L., Orrell, T., Nicolson, D., Bailly, N., Kirk, P., Bourgoin, T., DeWalt, R.E., Decock, W., De Wever, A., Nieukerken, E. van, Zarucchi, J. & Penev, L., eds. 2019. Species 2000 & ITIS Catalogue of Life. Published online. Accessed: 2019 Sept. 27. Reference page.
Govaerts, R. et al. 2019. Spergularia in Kew Science Plants of the World online. The Board of Trustees of the Royal Botanic Gardens, Kew. Published online. Accessed: 2019 Sept. 27. Reference page.
International Plant Names Index. 2019. Spergularia. Published online. Accessed: Sept. 27 2019. 2019. Spergularia. Missouri Botanical Garden. Published online. Accessed: 27 Sept. 2019.
USDA, ARS, Germplasm Resources Information Network. Spergularia in the Germplasm Resources Information Network (GRIN), U.S. Department of Agriculture Agricultural Research Service. Accessed: 09-Oct-10.

Vernacular names
Deutsch: Schuppenmieren
English: Sand Spurry
eesti: Sõlmhein
فارسی: زمین‌گستر
suomi: Solmukit
hornjoserbsce: Kołwróćik
magyar: Budavirág
lietuvių: Posmiltė
Nederlands: schijnspurrie
polski: Muchotrzew
русский: Торичник

Spergularia is a genus in the family Caryophyllaceae, containing salt-tolerant plants known as sandspurrys[1] and sea-spurreys. There are about 60 species.[2]
Selected species

Spergularia azorica – endemic to the archipelago of the Azores[3]
Spergularia atrosperma – blackseed sandspurry
Spergularia bocconei (Scheele) Graebn. – Boccone's sand-spurrey
Spergularia canadensis – Canadian sandspurry
Spergularia catalaunica Monnier
Spergularia diandra (Guss.) Boiss.
Spergularia echinosperma (Celak.) Asch. & Graebn.
Spergularia heldreichii Foucaud
Spergularia macrorrhiza (Loisel.) Heynh.
Spergularia macrotheca – sticky sandspurry
Spergularia marina (L.) Besser – lesser sea-spurrey
Spergularia media (L.) C.Presl – greater sea-spurrey
Spergularia nicaeensis Sarato ex Burnat
Spergularia platensis
Spergularia rubra (L.) J.Presl & C.Presl – red sand-spurrey
Spergularia rupicola Lebel ex Le Jol.
Spergularia salina J.Presl & C.Presl
Spergularia segetalis (L.) G.Don
Spergularia tasmanica
Spergularia tangerina P.Monnier
Spergularia villosa – hairy sandspurry

See also

Similar genera that have been taxonomically intertwined with Spergularia:


USDA, NRCS (n.d.). "Spergularia". The PLANTS Database ( Greensboro, North Carolina: National Plant Data Team. Retrieved 25 November 2015.
Flora of North America
Secretaria Regional do Ambiente e do Mar (2008), Plantas Endémicas dos Açores: Guia da Ilha do Faial, p.11; Flowering between May and September, the azorica is a small plant, sometimes woody at its base, and found in only strongly exposed coastal cliffs.

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