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Classification System: APG IV

Superregnum: Eukaryota
Regnum: Plantae
Cladus: Angiosperms
Cladus: Eudicots
Cladus: Core eudicots
Cladus: Asterids
Cladus: Campanulids
Ordo: Asterales

Familia: Asteraceae
Subfamilia: Asteroideae
Tribus: Astereae
Subtribus: Solidagininae
Genus: Tonestus
Species: T. eximius – T. graniticus – T. lyallii – T. pygmaeus

Tonestus A.Nelson, Bot. Gaz. 37: 262. (1904)

Type species: Tonestus lyallii A.Nelson, Bot. Gaz. 37: 262. (1904)


Nelson, A. 1904. Botanical Gazette; Paper of Botanical Notes 37: 262.
Brouillet, L., Urbatsch, L. & Roberts, R.P. 2004. Tonestus kingii and T. aberrans are related to Eurybia and the Machaerantherinae (Asteraceae: Astereae) based on nrDNA (ITS and ETS) data: reinstatement of Herrickia and a new genus, Triniteurybia. SIDA, Contributions to Botany 21(2): 889-900. JSTOR BHL Reference page.
Hassler, M. 2018. Tonestus. World Plants: Synonymic Checklists of the Vascular Plants of the World In: Roskovh, Y., Abucay, L., Orrell, T., Nicolson, D., Bailly, N., Kirk, P., Bourgoin, T., DeWalt, R.E., Decock, W., De Wever, A., Nieukerken, E. van, Zarucchi, J. & Penev, L., eds. 2018. Species 2000 & ITIS Catalogue of Life. Published online. Accessed: 2018 Nov. 21. Reference page.
International Plant Names Index. 2018. Tonestus. Published online. Accessed: Nov. 22 2018. 2018. Tonestus. Missouri Botanical Garden. Published online. Accessed: 22 Nov. 2018.

Tonestus, common name serpentweed,[2] is a genus of North American flowering plants in the aster family.[3][4][5][6][7][2][8]

The name is an anagram of the name of the related genus Stenotus.[3] Nestotus is similarly another anagram of the same name.[9]


Tonestus eximius (H.M.Hall) A.Nelson & J.F.Macbr. - CA NV
Tonestus graniticus (Tiehm & L.M.Shultz) G.L.Nesom & D.R.Morgan - NV
Tonestus lyallii (A.Gray) A.Nelson - Alberta, British Columbia, WA OR ID MT CO WY NV CA
Tonestus pygmaeus (Torr. & A.Gray) A.Nelson - MT WY CO NM

formerly included[1]

now in Herrickia Lorandersonia Toiyabea

Tonestus alpinus (L.C.Anderson & Goodrich) G.L.Nesom & D.R.Morgan - Toiyabea alpina (L.C.Anderson & Goodrich) R.P.Roberts, Urbatsch & Neubig
Tonestus kingii (D.C.Eaton) G.L.Nesom - Herrickia kingii (D.C.Eaton) Brouillet, Urbatsch & R.P.Roberts
Tonestus microcephalus (Cronquist) G.L.Nesom & D.R.Morgan - Lorandersonia microcephala (Cronquist) Urbatsch, R.P.Roberts & Neubig
Tonestus peirsonii (D.D.Keck) G.L.Nesom & D.R.Morgan - Lorandersonia peirsonii (D.D.Keck) Urbatsch, R.P.Roberts & Neubig


Flann, C (ed) 2009+ Global Compositae Checklist Archived 2014-12-03 at
Flora of North America
Nelson, Aven. 1904. Botanical Gazette 37(4): 262 in English
Botanic Garden and Botanical Museum Berlin-Dahlem. "Details for: Astereae". Euro+Med PlantBase. Freie Universität Berlin. Retrieved 2009-07-18.
UniProt. "Tribe Astereae". Retrieved 2009-07-18.
National Herbarium of New South Wales. "Genus Kippistia". New South Wales FloraOnline. Royal Botanic Gardens, Sydney. Retrieved 2009-07-18.
"Polyarrhena Cass". African Plants Database. South African National Biodiversity Institute, the Conservatoire et Jardin botaniques de la Ville de Genève and Tela Botanica. Retrieved 2009-07-18.
Tropicos, Tonestus A. Nelson
Roberts, Roland P., Urbatsch, Lowell Edward, & Neubig, Kurt Maximilian. 2005. Sida 21(3): 1647-1655
Biota of North America Program 2013 range distribution maps

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