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Cladus: Eukaryota
Regnum: Plantae
Divisio: Pteridophyta
Subdivisio: Pterophyta
Classis: Pteridopsida
Ordo: Polypodiales
Familia: Aspleniaceae
Genera: Antigramma - Asplenium - Camptosorus - Diellia - Holodictyum - Hymenasplenium - Loxoscaphe - Pleurosorus - Schaffneria - Sinephropteris - Thamnopteris

Vernacular names
English: spleenworts
Magyar: Fodorkafélék
Nederlands: Aspleniumachtigen
日本語: チャセンシダ科
Русский: Костенцовые


Aspleniaceae A.B.Frank


* Smith, A.R. et al. (2006). A classification for extant ferns. Taxon 55 (3): 705–731 pdf
* Checklist of ferns and fern allies [1].


The Aspleniaceae (spleenworts) is a family of ferns, included in the order Polypodiales or in some classifications as the only family in the order Aspleniales.

Members of the family all have intramarginal, linear sori with a flap-like indusium arising along one edge. Most pteridologists today consider this family of consisting of just one to three genera. Others still maintain segregate genera such as Phyllitis and Ceterach; however, the species segregated into these genera all hybridize readily with undisputed Asplenium species.[specify] A recent phylogenenetic study of Aspleniaceae (Murukami et al. 1999) shows that species segregated as Camptosorus and Neottopteris are nested within Asplenium and recommends that they be included in that genus, but suggests that Hymenasplenium (including Boniniella) and Phyllitis are distantly related to other Asplenium species and should be recognized at the generic level.

The genus Diellia, consisting of six species found only in Hawaii, was long considered to be independent, but now has been shown to nest within Asplenium.[1]


1. ^ Schneider H et al. (2005 Feb 22), "Origin of the endemic fern genus Diellia coincides with the renewal of Hawaiian terrestrial life in the Miocene", Proc Biol Sci. 272 (1561): 455–60, PMID 15734701

* Germplasm Resources Information Network: Aspleniaceae
* Murakami, N., S. Nogami, M. Watanabe, K. Iwatsuki. 1999. Phylogeny of Aspleniaceae inferred from rbcL nucleotide sequences. American Fern Journal 89: 232-243. doi:10.2307/1547233

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Source: Wikispecies, Wikipedia: All text is available under the terms of the GNU Free Documentation License