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Superregnum: Eukaryota
Regnum: Plantae
Divisio: Tracheophyta
Divisio: Pteridophyta
Classis: Polypodiopsida
Ordo: Polypodiales

Familia: Cystopteridaceae
Genera: Acystopteris – Cystoathyrium – Cystopteris – Gymnocarpium

Cystopteridaceae Schmakov (2001)

Type genus: Cystopteris Bernh.


Cystopteridinae Payer, Bot. Crypt.: 198 (1850).


Shmakov, A. I. (2001). Turczaninowia 4 (1-2): 60.
Christenhusz, M.J.M., Zhang, X.-C. & Schneider, H. 2011. A linear sequence of extant families and genera of lycophytes and ferns. Phytotaxa 19: 7–54. DOI: 10.11646/phytotaxa.19.1.2 Open access. Reference page

Vernacular names
čeština: Puchýřníkovité
فارسی: بادکنکی‌سرخسان
suomi: Loikkokasvit
한국어: 한들고사리과
polski: Paprotnicowate
русский: Пузырниковые
українська: Міхурницеві

Cystopteridaceae is a family of ferns in the order Polypodiales. In the Pteridophyte Phylogeny Group classification of 2016 (PPG I), the family is placed in the suborder Aspleniineae of the order Polypodiales, and includes three genera.[2] Alternatively, it may be treated as the subfamily Cystopteridoideae of a very broadly defined family Aspleniaceae.[3]

Cystopteridaceae are small or medium-sized ferns in forests and crevices.[4] They generally have thin laminae, and small, round, naked sori.

Three genera are accepted in the PPG I classification, and by the Checklist of Ferns and Lycophytes of the World as of November 2019:[2][1]

Acystopteris Nakai
Cystopteris Bernh.
Gymnocarpium Newman

Cystopteris chinensis has been placed in a separate genus, Cystoathyrium, but the genus is not accepted by recent sources.[5][6]
Phylogenetic relationships

The following diagram for the Aspleniineae (eupolypods II), based on Lehtonen, 2011,[7] and Rothfels & al., 2012,[8] shows a likely phylogenic relationship between the Cystopteridaceae and the other families of the clade.

Aspleniineae (eupolypods II)











Gymnocarpium and Cystopteris have been found to be sister to the remaining Aspleniineae.[4]

Hassler, Michael & Schmitt, Bernd (November 2019). "Cystopteridaceae". Checklist of Ferns and Lycophytes of the World. 8.11. Retrieved 2019-11-27.
PPG I (2016). "A community-derived classification for extant lycophytes and ferns". Journal of Systematics and Evolution. 54 (6): 563–603. doi:10.1111/jse.12229. S2CID 39980610.
Christenhusz, Maarten J.M. & Chase, Mark W. (2014). "Trends and concepts in fern classification". Annals of Botany. 113 (9): 571–594. doi:10.1093/aob/mct299. PMC 3936591. PMID 24532607.
Maarten J. M. Christenhusz; Xian-Chun Zhang; Harald Schneider (2011). "A linear sequence of extant families and genera of lycophytes and ferns" (PDF). Phytotaxa. 19: 7–54. doi:10.11646/phytotaxa.19.1.2.
Hassler, Michael & Schmitt, Bernd (November 2019). "Cystoathyrium chinense". Checklist of Ferns and Lycophytes of the World. 8.11. Retrieved 2019-11-27.
"Cystopteris chinensis Ching". Plants of the World Online. Royal Botanic Gardens, Kew. Retrieved 2019-11-27.
Samuli Lehtonen (2011). "Towards Resolving the Complete Fern Tree of Life" (PDF). PLOS ONE. 6 (10): e24851. doi:10.1371/journal.pone.0024851. PMC 3192703. PMID 22022365.
Carl J. Rothfels; Anders Larsson; Li-Yaung Kuo; Petra Korall; Wen- Liang Chiou; Kathleen M. Pryer (2012). "Overcoming Deep Roots, Fast Rates, and Short Internodes to Resolve the Ancient Rapid Radiation of Eupolypod II Ferns". Systematic Biology. 61 (1): 490–509. doi:10.1093/sysbio/sys001. PMID 22223449.

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