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Classification System: APG IV

Superregnum: Eukaryota
Regnum: Plantae
Cladus: Angiosperms
Cladus: Eudicots
Cladus: Core eudicots
Cladus: Asterids
Cladus: Campanulids
Ordo: Asterales

Familia: Asteraceae
Subfamilia: Carduoideae
Tribus: Cardueae
Subtribus: Carduinae
Genus: Cousinia
Sectiones: C. sect. Acanthotoma – C. sect. Actinia – C. sect. Albidae – C. sect. Albolinia – C. sect. Alpinae – C. sect. Argenteae – C. sect. Badghysia – C. sect. Carduncellus – C. sect. Chaetocephalae – C. sect. Chaetolepides – C. sect. Chrysantha – C. sect. Chrysoptera – C. sect. Congestae – C. sect. Coronophora – C. sect. Cousinia – C. sect. Ctenarctium – C. sect. Cynaroideae – C. sect. Decumbentes – C. sect. Dichotomae – C. sect. Eriocousinia – C. sect. Fragillimae – C. sect. Glaphyrocephalae – C. sect. Harazensis – C. sect. Haussknechtianae – C. sect. Helianthae – C. sect. Hemsleyanae – C. sect. Homalochaete – C. sect. Immitantes – C. sect. Jurineopsis – C. sect. Kopetdaghia – C. sect. Kuekenthalia – C. sect. Lachnosphaerae – C. sect. Lasiandrae – C. sect. Leiocaules – C. sect. Lepidae – C. sect. Leucocaulon – C. sect. Lopholepis – C. sect. Macilenta – C. sect. Microcarpae – C. sect. Molles – C. sect. Myriotomae – C. sect. Nonarctium – C. sect. Neurocentrae – C. sect. Olgaeanthe – C. sect. Oliganthe – C. sect. Platyacanthae – C. sect. Pseudactinia – C. sect. Pugioniferae – C. sect. Pulchellae – C. sect. Rigidissima – C. sect. Racemosae – C. sect. Scariosae – C. sect. Sciadocousinia – C. sect. Serratuloides – C. sect. Singulares – C. sect. Sphaerocephalae – C. sect. Spinuliferae – C. sect. Stenocephalae – C. sect. Stenoloma – C. sect. Stocksianae – C. sect. Tenellae – C. sect. Tricolorae – C. sect. Winklerianae

Overview of species
a

C. abbreviata – C. acanthoblephara – C. acanthodendron – C. acanthophysa – C. acanthostephana – C. acrodroma – C. actinacantha – C. actinia – C. adenophora – C. adenostegia – C. adenosticta – C. adnata – C. affinis – C. afghanica – C. agelocephala – C. agridaghensis – C. aggregata – C. aintabensis – C. aitchisonii – C. alaica – C. alata – C. albertoregelia – C. albescens – C. albicaulis – C. albida – C. albiflora – C. alepideae – C. aleppica – C. alexeenkoana – C. alfredii – C. algurdina – C. aligudarzensis – C. allolepis – C. alpestris – C. alpicola – C. ambigens – C. ammophila – C. amoena – C. andarabensis – C. androssovii – C. angreni – C. annua – C. anoplophylla – C. antonowii – C. apiculata – C. aptera – C. arachnoidea – C. arakensis – C. araneosa – C. araneotexta – C. araxena – C. archibaldii – C. arctotidifolia – C. ardalensis – C. arenaria – C. argentea – C. ariana – C. arida – C. armena – C. aspera – C. assadii – C. assyriaca – C. astracanica – C. atripurpurea – C. atrobracteata – C. atropatana – C. attariae – C. aucheri – C. auriculata – C. austrojordanica – C. autrani – C. azerbaidjanica – C. azmarensis
b

C. baberi – C. bachtiarica – C. badhysi – C. balchanica – C. baluchistanica – C. bamianica – C. baranovii – C. barbeyi – C. barezica – C. batalinii – C. bazoftensis – C. beauverdiana – C. beckeri – C. belangeri – C. bicolor – C. bienerti – C. bijarensis – C. bipinnata – C. birandiana – C. blepharobasis – C. bobekii – C. bobrovii – C. boissieri – C. bonvalotii – C. bornmuelleri – C. botschantzevii – C. brachyptera – C. brevicaulis – C. bucharica – C. bungeana – C. buphthalmoides – C. butkovii
c

C. caesarea – C. caespitosa – C. calcitrapa – C. calcitrapella – C. calocephala – C. calolepis – C. calva – C. campyloraphis – C. cana – C. candicans – C. candolleana – C. canescens – C. carduchorum – C. carduncelloidea – C. caroli-henrici – C. carthamoides – C. cataonica – C. catenata – C. cavarae – C. centauroides – C. ceratophora – C. chaetocephala – C. chamaepeuce – C. chamaepeucides – C. chejrabadensis – C. chionophila – C. chitralensis – C. chlorocephala – C. chlorosphaera – C. chlorothyrsa – C. chrysacantha – C. chrysandra – C. chrysantha – C. chrysochlora – C. cirsioides – C. cisdarvasica – C. coerulea – C. commutata – C. concinna – C. concolor – C. congesta – C. consanguinea – C. contumax – C. cordifolia – C. coronata – C. corymbosa – C. crassipes – C. crispa – C. cryptadena – C. curvibracteata – C. cylindracea – C. cylindrocephala – C. cymbolepis – C. cynaroides – C. czatkalica – C. czerniakowskae
d

C. dalahuensis – C. daralaghezica – C. darwasica – C. dasylepis – C. davisiana – C. decipiens – C. decolorans – C. decumbens – C. decurrens – C. decurrentifolia – C. denaensis – C. deserti – C. dichotoma – C. dichromata – C. diezii – C. dimoana – C. dipterocarpa – C. discolor – C. dissecta – C. dissectifolia – C. divaricata – C. dolichoclada – C. dolicholepsis – C. dshisakensis – C. dubia
e

C. eburnea – C. ecbatanensis – C. echinocephala – C. edmondsonii – C. egens – C. elata – C. elburzensis – C. eleonorae – C. elephantina – C. elwendensis – C. erectispina – C. erinacea – C. eriobasis – C. eriophylla – C. eriorrhiza – C. eriotricha – C. erivanensis – C. ermenekensis – C. eryngioides – C. esfandiarii – C. euchlora – C. eugenii – C. euphratica – C. eurylepis
f

C. fabrorum – C. falcinella – C. falconeri – C. fallax – C. farsistanica – C. fascicularis – C. fedorovii – C. ferghanensis – C. ferruginea – C. fetissowii – C. finitima – C. firuzkuhensis – C. foliosa – C. fragilis – C. fragillima – C. franchetii – C. fraternella – C. freitagii – C. freynii – C. fursei
g

C. gabrielae – C. gaharensis – C. gatchsaranica – C. gaubae – C. gedrosiaca – C. ghahremanii – C. ghorana – C. gigantolepis – C. gigantoptera – C. gigantosphaera – C. gilanica – C. gilliatii – C. glabriseta – C. glandulosa – C. glaphyra – C. glaphyrocephala – C. glaucopsis – C. glochidiata – C. gmelini – C. gnezdilloi – C. gomolitzkii – C. gontscharowii – C. gracilis – C. graminifolia – C. grandiceps – C. grandis – C. grantii – C. greuteri – C. griffithiana – C. grigoriewii – C. grisea – C. gulczensis – C. gymnoclada
h

C. hablitzii – C. haeckeliae – C. hakkarica – C. halysensis – C. hamadae – C. hamadanensis – C. hamosa – C. harazensis – C. hastifolia – C. haussknechtii – C. hazarensis – C. hedgei – C. heliantha – C. hemsleyana – C. hergtiana – C. heterogeneta – C. heteroloba – C. heteromorpha – C. heterophylla – C. hilariae – C. hohenackeri – C. hololeuca – C. hoplites – C. hoplophylla – C. horrida – C. humilis – C. hybrida – C. hymenostephanus – C. hypochionea – C. hypoleuca – C. hystricocephala – C. hystrix
i

C. iconica – C. ilicifolia – C. iljinii – C. immitans – C. immitantiformis – C. incompta – C. inflata – C. infundibularis – C. insignis – C. integrifolia – C. intermedia – C. intertexta – C. iranshahriana – C. iranshahrii – C. irritans – C. isfahanica – C. iskanderi
j

C. jacobsii – C. jassyensis – C. juzepczukii
k

C. kaluensis – C. kamarbandensis – C. karkasensis – C. kasachanstanica – C. kataghanica – C. kazachorum – C. keredjensis – C. kermanensis – C. kermanshahensis – C. kerstanii – C. khashensis – C. khorasanica – C. khorramabadensis – C. kilouyensis – C. kirrindica – C. knorringiae – C. koelzii – C. kokanica – C. komarowii – C. komidjanensis – C. kopi-karadaghensis – C. kornhuberi – C. korowiakowi – C. kotschyi – C. kovalevskiana – C. krauseana – C. kuekenthalii – C. kuramensis – C. kurdistanica – C. kurubasgecidiensis
l

C. lachnobasis – C. lachnosphaera – C. lactiflora – C. laetevirens – C. lanata – C. lancifolia – C. lasiandra – C. lasiolepis – C. lasiophylla – C. leatherdalei – C. leiocephala – C. lepida – C. leptacma – C. leptocampyla – C. leptocephala – C. leptoclada – C. leptocladoides – C. leptolepis – C. leptomera – C. leucantha – C. libanotica – C. lignosissima – C. linczewskii – C. litvinovii – C. lomakinii – C. longibracteata – C. longifolia – C. lordeganensis – C. lucida – C. lurestanica – C. lurorum – C. lyrata
m

C. maasoumii – C. macrocephala – C. macrolepis – C. macroptera – C. magnifica – C. malacophylla – C. malurensis – C. manouchehrii – C. maracandica – C. margaritae – C. margiana – C. mattfeldii – C. mazu-shirinensis – C. megalomastix – C. meghrica – C. mehreganii – C. meshhedensis – C. mesomorpha – C. microcarpa – C. microcephala – C. millefontana – C. mindshelkensis – C. minkwitziae – C. miserabilis – C. moabitica – C. mobayenii – C. mogensii – C. mogoltavica – C. mollis – C. monocephala – C. mozaffarianii – C. mozdouranensis – C. mucida – C. mulgediifolia – C. multiloba – C. murgabica – C. mutehensis – C. myrioglochis – C. myriolepis – C. myriotoma
n

C. nabelekii – C. naria – C. neglecta – C. neurocentra – C. newesskyana – C. ninae – C. nischapurensis – C. nivea – C. niveocoronata – C. noeana – C. novissima – C. nujianensis
o

C. odontolepis – C. olgae – C. oligocephala – C. olivieri – C. omissa – C. omphalodes – C. onopordioides – C. onopordon – C. oophora – C. oopoda – C. oreodoxa – C. oreoxerophila – C. orientalis – C. orthacantha – C. orthoclada – C. ortholepis – C. orthoneura – C. oshtorankuhensis – C. ottonis – C. outichaschensis – C. ovczinnikovii – C. oxiana – C. oxytoma
p

C. panjshirensis – C. pannosa – C. pannosiformis – C. papillosa – C. paraatripurpura – C. parjumanensis – C. parsana – C. parviceps – C. parviziana – C. patentispina – C. pauciramosa – C. pectinata – C. pedunculata – C. peduncularis – C. pergamacea – C. perovskiensis – C. persarum – C. persica – C. perspolitana – C. pestalozzae – C. phyllocephala – C. pichleriana – C. pinarocephala – C. pineticola – C. piptocephala – C. platyacantha – C. platylepis – C. platyptera – C. platyraphis – C. platystegia – C. podlechii – C. podophylla – C. polycephala – C. polyneura – C. polytimetica – C. porphyrochrysea – C. porphyrostephana – C. postiana – C. praestans – C. prasina – C. princeps – C. prolifera – C. proalotiixima – C. psammophila – C. pseudactinia – C. pseudoaffinis – C. pseudarcticum – C. pseudobonvalotii – C. pseudocandolleana – C. pseudocirsium – C. pseudodshisakensis – C. pseudolanata – C. pseudomollis – C. pseudostenolepis – C. pterocarpa – C. pterocaulos – C. pugionifera – C. pulchella – C. pulcherantha – C. pulchra – C. pulvinaris – C. pungens – C. purpurea – C. pusilla – C. pycnocephala – C. pycnoloba – C. pygmaea
q

C. qaisarensis – C. qandilica – C. qaradaghensis – C. qarehbilensis – C. quettensis
r

C. racemosa – C. raddeana – C. radians – C. ramosissima – C. ramulosa – C. raphiostegia – C. rava – C. rawanduzensis – C. rechingerae – C. rechingerorum – C. recurvata – C. regelii – C. renominata – C. resinosa – C. rhabdodes – C. rhaphiocephala – C. rhodantha – C. rhombiformis – C. rigida – C. rigidissima – C. robusta – C. rosea – C. rotundifolia – C. rubiginosa – C. rudis – C. rufidola
s

C. sabalanica – C. sabzevarensis – C. sagittata – C. sahandica – C. sakawensis – C. salangensis – C. saloukensis – C. sanandajensis – C. sarawschanica – C. sardashtensis – C. satdagensis – C. scabrida – C. scala – C. scariosa – C. scheibeana – C. schepsaica – C. schindleriana – C. schiraziaca – C. schiraziana – C. schischkinii – C. schistoptera – C. schistosa – C. schtschurowskiana – C. schlagintweiti – C. schmalhausenii – C. schtschurowskiana – C. schugnanica – C. scleracantha – C. sclerolepis – C. sclerophylla – C. sefidiana – C. seidlitzii – C. semidecurrens – C. semilacera – C. serratuloides – C. severtzovii – C. shahrestanica – C. shahvarica – C. sharifii – C. sheidaii – C. shibarensis – C. shorlughensis – C. shugnanica – C. shulabadensis – C. sicigera – C. silvanica – C. silyboides – C. simulatrix – C. singularis – C. sinjarensis – C. sintenisii – C. sivasica – C. smirnowii – C. sogdiana – C. sororia – C. spathulata – C. speciosa – C. sphaerocephala – C. spiridonovii – C. splendida – C. sporadocephala – C. spryginii – C. stahliana – C. stapfiana – C. stechmanniopsis – C. stellaris – C. stenocalathia – C. stenocephala – C. stenophylla – C. stephanophora – C. stereolepis – C. stereoneura – C. stocksii – C. straussii – C. stricta – C. strobilocephala – C. stroterolepis – C. subappendiculata – C. subcandicans – C. subinflata – C. submutica – C. subpectinata – C. subscaposa – C. subtilis – C. sylvicola – C. syrdariensis
t

C. tabrisiana – C. takharensis – C. talassica – C. tamarae – C. tashkurghanica – C. taybadensis – C. tedshenica – C. tenella – C. tenuifolia – C. tenuiramula – C. tenuisecta – C. tenuispina – C. termei – C. tetanocephala – C. thamnodes – C. thomsonii – C. tianschanica – C. tirinensis – C. tomentella – C. trachylepis – C. trachyphylla – C. trachyphyllaria – C. trachycanthoides – C. transiliensis – C. transoxana – C. triacantha – C. triceps – C. trichophora – C. tricolor – C. trollii – C. tscherneviae – C. turcomanica – C. turkestanica – C. turkmenorum
u

C. ulotoma – C. umbilicata – C. unaiensis – C. urumiensis
v

C. verbascifolia – C. verticillaris – C. vicaria – C. volkii – C. vvedenskyi
w

C. waldheimiana – C. wendelboi – C. wesheni – C. wheeler-hainesii – C. wilhelminae – C. winkleriana – C. woronowi
x

C. xanthacantha – C. xanthina – C. xanthiocephala – C. xanthothyrsa – C. xanthula – C. xiphiolepis
y

C. yasujensis
z

C. zagrica

Nothospecies

C. × kurubasgecidiensis
Source(s) of checklist:
Name

Cousinia Cass. in F. Cuvier, Dict. Sci. Nat., ed. 2. 47: 503. 1827.

Type species: Cousinia carduiformis Cass. (1827), synonym of Cousinia orientalis (Adams) C.Koch (1851)

Synonyms

Heterotypic
Tiarocarpus Rech.f., Anz. Österr. Akad. Wiss., Math.-Naturwiss. Kl. 108: 5. 1972.
Lipskyella Juz., Trudy Bot. Inst. Akad. Nauk S.S.S.R., Ser. 1, Fl. Sist. Vyssh. Rast. 3: 323. 1937.

References
Primary references

Cassini, A.H.G. de 1827. Saussurée [avec tableau méthodique des genres et sous-genres composant la tribu des Carlinées] in: Cuvier, G.-F. 1827. Dictionnaire des Sciences Naturelles, dans lequel on traite méthodiquement des différens êtres de la nature, considérés soit en eux-mêmes, d'aprés l'état actuel de nos connoissances, soit relativement à l'utilité quén peuvent retirer la médecine, l'agriculture, le commerce et les arts. Edition 2 47: 494–513 (!503).

Additional references

Sennikov, A.N. 2010. A revision of Cousinia sections Alpinae (syn. Carduncellus), Subappendiculatae and Tianschanicae (Asteraceae) in the Kirghizian Tian-Shan and the neighbouring territories. Phytotaxa 5: 1–30. DOI: 10.11646/phytotaxa.5.1.1 Paywall Reference page.
López-Vinyallonga, S., Romaschenko, K., Susanna, A. & Garcia-Jacas, N. 2011. Systematics of the arctioid group: disentangling Arctium and Cousinia (Cardueae, Carduinae). Taxon 60(2): 539–554. DOI: 10.1002/tax.602020 JSTORPDF Reference page.
Ahmad, S.A., Rastegar, A. & Attar, F. 2017. Cousinia azmarensis (Asteraceae, Cardueae), a New Species from Kurdistan, Iraq. Harvard Papers in Botany 22(1): 71–73. DOI: 10.3100/hpib.v22iss1.2017.n8Paywall Reference page.

Links

Hassler, M. 2020. Cousinia. World Plants: Synonymic Checklists of the Vascular Plants of the World In: Roskovh, Y., Abucay, L., Orrell, T., Nicolson, D., Bailly, N., Kirk, P., Bourgoin, T., DeWalt, R.E., Decock, W., De Wever, A., Nieukerken, E. van, Zarucchi, J. & Penev, L., eds. 2020. Species 2000 & ITIS Catalogue of Life. Published on the internet. Accessed: 2020 Jan. 15. Reference page.
Govaerts, R. et al. 2020. Cousinia in Kew Science Plants of the World online. The Board of Trustees of the Royal Botanic Gardens, Kew. Published on the internet. Accessed: 2020 Jan. 15. Reference page.
International Plant Names Index. 2020. Cousinia. Published online. Accessed: Jan. 15 2020.
Tropicos.org 2020. Cousinia. Missouri Botanical Garden. Published on the internet. Accessed: 2020 Feb 22.

The genus Cousinia of the tribe Cardueae (= Cynareae) is in its current circumscription one of the larger genera in the Asteraceae, with approximately 650-700 species distributed in central and western Asia.[1][2]

Many of the species in this genus were once classified in genus Arctium (the burdocks).
Species
Main article: List of Cousinia species
References

López-Vinyallonga, Sara; Romaschenko, Kostyantyn; Susanna, Alfonso; Garcia-Jacas, Núria (2011). "Systematics of the Arctioid group: Disentangling Arctium and Cousinia (Cardueae, Carduinae)". Taxon. 60 (2): 539–554. doi:10.1002/tax.602020. hdl:10261/35343.
"Cousinia Cass". Plants of the World Online. Board of Trustees of the Royal Botanic Gardens, Kew. 2021. Retrieved 2 September 2021.

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