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Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Superphylum: Deuterostomia
Phylum: Chordata
Subphylum: Vertebrata
Infraphylum: Gnathostomata
Megaclassis: Osteichthyes
Cladus: Sarcopterygii
Cladus: Rhipidistia
Cladus: Tetrapodomorpha
Cladus: Eotetrapodiformes
Cladus: Elpistostegalia
Superclassis: Tetrapoda
Cladus: Reptiliomorpha
Cladus: Amniota
Classis: Reptilia
Cladus: Eureptilia
Cladus: Romeriida
Subclassis: Diapsida
Cladus: Sauria
Infraclassis: Lepidosauromorpha
Superordo: Lepidosauria
Ordo: Squamata
Cladus: Unidentata Episquamata

Subordo: Lacertoidea
Infraordo: Lacertibaenia

Familia: Lacertidae
Subfamilia: Lacertinae
Genus: Heliobolus
Species (6): H. bivari – H. crawfordi – H. lugubris – H. neumanni – H. nitidus – H. spekii

Heliobolus Fitzinger, 1843: 20

Type species: Eremias lugubris Duméril & Bibron, 1839, in error [=Lacerta lugubris Smith, 1838], by original designation.

Primary references

Fitzinger, L. 1843. Systema Reptilium. Fasciculus primus. Amblyglossae. Braumüller et Seidel: Wien. vi + 106 pp. BHL Reference page.

Additional references

Marques, M.P., Ceríaco, L.M.P., Heinicke, M.P., Chehouri, R.M., Conradie, W., Tolley, K.A. & Bauer, A.M. 2022. The Angolan bushveld lizards, genus Heliobolus Fitzinger, 1843 (Squamata: Lacertidae): Integrative taxonomy and the description of two new species. Vertebrate Zoology 72: 745–769. DOI: 10.3897/vz.72.e85269 Open access Reference page.


Uetz, P. & Hallermann, J. 2023. Heliobolus . The Reptile Database. Accessed on 30 March 2022.

Vernacular names
English: Sand Lizards

Heliobolus is a genus of lizards of the family Lacertidae. The genus is endemic to Africa.[1]

There are six recognized species:[1][2]

Heliobolus bivari Marques, Ceríaco, Heinicke, Chehouri, Conradie, Tolley, & Bauer, 2022 – Bivar’s bushveld lizard
Heliobolus crawfordi Marques, Ceríaco, Heinicke, Chehouri, Conradie, Tolley, & Bauer, 2022 – Crawford-Cabral’s bushveld lizard
Heliobolus lugubris (A. Smith, 1838) – bushveld lizard, mourning racerunner, black and yellow sand lizard
Heliobolus neumanni (Tornier, 1905) – Neumann's sand lizard
Heliobolus nitidus (Günther, 1872) – glittering sand lizard
Heliobolus spekii (Günther, 1872) – Speke's sand lizard

Nota bene: A binomial authority in parentheses indicates that the species was originally described in a genus other than Heliobolus.

Heliobolus at the Reptile Database. Accessed 28 March 2020.

Genus Heliobolus. The Taxonomicon.

Further reading

Branch, Bill (2004). Field Guide to Snakes and other Reptiles of Southern Africa. Third Revised edition, Second impression. Sanibel Island, Florida: Ralph Curtis Books. 399 pp. ISBN 0-88359-042-5. (Genus Heliobolus, p. 161).
Fitzinger L (1843). Systema Reptilium, Fasciculus Primus, Amblyglossae. Vienna: Braumüller & Seidel. 106 pp. + indices. (Heliobolus, new genus, p. 20). (in Latin).

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