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Classification System: APG IV

Superregnum: Eukaryota
Regnum: Plantae
Cladus: Angiosperms
Cladus: Eudicots
Cladus: Core eudicots
Ordo: Caryophyllales

Familia: Nyctaginaceae
Tribus: Nyctagineae
Genus: Anulocaulis
Species: A. annulatusA. eriosolenus – A. hintoniorum – A. leiosolenusA. reflexus


Anulocaulis Standl. Contr. U.S. Natl. Herb. 12: 374. (1909)

Type species: Anulocaulis eriosolenus (A. Gray) Standl. Contr. U.S. Natl. Herb. 12: 375. (1909)


Standley, P.C. 1909. Contributions from the United States National Herbarium 12(8): 374–375.
Hassler, M. 2017. Anulocaulis. World Plants: Synonymic Checklists of the Vascular Plants of the World In: Roskovh, Y., Abucay, L., Orrell, T., Nicolson, D., Bailly, N., Kirk, P., Bourgoin, T., DeWalt, R.E., Decock, W., De Wever, A., Nieukerken, E. van, Zarucchi, J. & Penev, L., eds. 2017. Species 2000 & ITIS Catalogue of Life. Published online. Accessed: 2017 May 12. Reference page.
Hernández-Ledesma, P., Olvera, H.F. & Ochoterena, H. 2010. Cladistic Analysis and Taxonomic Synopsis of Anulocaulis (Nyctaginaceae) Based on Morphological Data. Systematic Botany 35(4): 858-876. DOI: 10.1600/036364410X539916 Reference page.
International Plant Names Index. 2017. Anulocaulis. Published online. Accessed: 12 May 2017. 2017. Anulocaulis. Missouri Botanical Garden. Published online. Accessed: 12 May 2017.

Vernacular names
English: Ringstems

Anulocaulis is a small genus of flowering plants known generally as ringstems. These five species are sometimes treated as members of genus Boerhavia. Ringstems are thickly-rooted perennial wildflowers with glutinous brown bands at their stem internodes, the trait which gives them their common and Latin names. They bear tubular flowers at the tops of their stems. Ringstems are native to North America.


Anulocaulis annulatus - valley ringstem
Anulocaulis eriosolenus - Big Bend ringstem
Anulocaulis gypsogenus - gypsum ringstem
Anulocaulis leiosolenus - southwestern ringstem
Anulocaulis reflexus - Texas ringstem[1]

See also



"Species in Anulocaulis".

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