Cephalanthus occidentalis

Classification System: APG IV

Superregnum: Eukaryota
Regnum: Plantae
Cladus: Angiosperms
Cladus: Eudicots
Cladus: Core eudicots
Cladus: Asterids
Cladus: Lamiids
Ordo: Gentianales

Familia: Rubiaceae
Subfamilia: Cinchonoideae
Tribus: Naucleeae
Genus: Cephalanthus
Species: Cephalanthus occidentalis
Name

Cephalanthus occidentalis L.
References

Linnaeus, C. 1753.
Linnaeus, C. 1753. Species Plantarum. Tomus I: 5. Reference page. *
USDA, ARS, Germplasm Resources Information Network. Cephalanthus occidentalis in the Germplasm Resources Information Network (GRIN), U.S. Department of Agriculture Agricultural Research Service. Accessed: 07-Oct-06.

Vernacular names
čeština: hlavoš západní
English: buttonbush
slovenčina: hlavoš západný

Cephalanthus occidentalis is a species of flowering plant in the family Rubiaceae, that is native to eastern and southern North America. Common names include buttonbush, common buttonbush, button-willow, buck brush, and honey-bells.

Description

Cephalanthus occidentalis is a deciduous shrub or small tree that averages 1–3 m (3.3–9.8 ft) in height, but can reach 6 m (20 ft). The leaves are opposite or in whorls of three, elliptic to ovate, 7–18 cm (2.8–7.1 in) long and 4–10 cm (1.6–3.9 in) broad, with a smooth edge and a short petiole. The flowers are arranged in a dense spherical inflorescence 2–3.5 cm (0.79–1.38 in) in diameter on a short peduncle. Each flower has a fused white to pale yellow four-lobed corolla forming a long slender tube connecting to the sepals. The stigma protrudes slightly from the corolla. The fruit is a spherical cluster of achenes (nutlets).[4]
Taxonomy

There are two varieties, not considered distinct by all authorities:

Cephalanthus occidentalis var. occidentalis (syn. var. pubescens) – common buttonbush. Eastern North America from Nova Scotia west to Minnesota and south to Florida and east Texas.
Cephalanthus occidentalis var. californicus – California button-willow. Southwestern North America, from west Texas west to California (Sierra Nevada foothills, San Joaquin Valley, Sacramento Valley, and the Inner North Coast Ranges) and south to Mexico and Central America.

Habitat

Buttonbush is a common shrub of many wetland habitats in its range, including swamps, floodplains, mangrove, pocosin, riparian zones, and moist forest understory.[5] It is a member of the flora in the Everglades.[5]
Ecology

Waterfowl and other birds eat the seeds. Wood ducks utilize the plant as nest protection, and mallards eat the fruit.[6] Deer browse the foliage, which is poisonous to livestock.[7] Insects and hummingbirds take the nectar, with bees using it to make honey.[5][8] It is a larval host to the hydrangea sphinx, the royal walnut moth, and the titan sphinx.[9]
Distribution
In East Texas

The species occurs in eastern North America with disjunct populations occurring in the west. In Canada, it occurs from southern Ontario and Quebec east to New Brunswick and south-western Nova Scotia. Besides the eastern United States, and eastern regions of the Midwest, notable areas range into Arizona, the Mogollon Rim, and other mountain ranges; in California, the entire San Joaquin Valley[10] West of the Great Plains and the Rocky Mountains, only western Texas, Arizona, and California find C. occidentalis.
Uses
Medicinal

Cephalanthus occidentalis has a number of historical medicinal uses, but it is also toxic due to the presence of cephalanthin.[5][8]
Cultivation

Buttonbush is cultivated as an ornamental plant for a nectar source or 'honey plant' and for aesthetics in gardens and native plant landscapes, and is planted on slopes to help control erosion.[11] Buttonbush is a suitable shrub for butterfly gardens.
San Joaquin Valley landmark tree
The Buttonwillow Tree in Buttonwillow, California

The town of Buttonwillow, California was named for the buttonbush (Cephalanthus occidentalis). A lone buttonbush served as a landmark on an old trans-San Joaquin Valley trail, and was used by ancient Yokut Indians as a meeting place. It later became the site of settlers' stock rodeos. This buttonbush tree is listed as California Historical Landmark No. 492, and is now known as the "Buttonwillow Tree."[12]
References

Maiz-Tome, L. (2016). "Cephalanthus occidentalis". IUCN Red List of Threatened Species. 2016: e.T64310261A67729125. doi:10.2305/IUCN.UK.2016-1.RLTS.T64310261A67729125.en. Retrieved 6 May 2020.
"Cephalanthus occidentalis". Germplasm Resources Information Network (GRIN). Agricultural Research Service (ARS), United States Department of Agriculture (USDA). Retrieved 2010-01-04.
"Cephalanthus occidentalis" (PDF). Digital Representations of Tree Species Range Maps from "Atlas of United States Trees" by Elbert L. Little, Jr. (and other publications). United States Geological Survey.
"Cephalanthus occidentalis L. buttonbush" (PDF). Wildland Shrubs of the United States and its Territories: Thamnic Descriptions. United States Forest Service. Retrieved 2009-09-14.
"Cephalanthus occidentalis". Fire Effects Information System. United States Forest Service.
Niering, William A.; Olmstead, Nancy C. (1985) [1979]. The Audubon Society Field Guide to North American Wildflowers, Eastern Region. Knopf. p. 764. ISBN 0-394-50432-1.
Little, Elbert L. (1980). The Audubon Society Field Guide to North American Trees: Eastern Region. New York: Knopf. p. 667. ISBN 0-394-50760-6.
"Common Buttonbush Cephalanthus occidentalis L." (PDF). Natural Resources Conservation Service Plant Guide. United States Department of Agriculture.
The Xerces Society (2016), Gardening for Butterflies: How You Can Attract and Protect Beautiful, Beneficial Insects, Timber Press.
Little Jr., Elbert L. (1976). "Map 34, Cephalanthus occidentalis". Atlas of United States Trees. Vol. 3 (Minor Western Hardwoods). US Government Printing Office. LCCN 79-653298. OCLC 4053799.
O'Sullivan, Penelope (2007). The Homeowner's Complete Tree & Shrub Handbook: The Essential Guide to Choosing, Planting, and Maintaining Perfect Landscape Plants. Storey Publishing. p. 197. ISBN 978-1-58017-571-5.

"CHL # 492 Buttonwillow Tree Kern". www.californiahistoricallandmarks.com.

USDA Plants Profile: Cephalanthus occidentalis
Missouriplants: Cephalanthus occidentalis var. occidentalis
Jepson Flora Project: Cephalanthus occidentalis var. californicus