Gratioleae

Life-forms

Classification System: APG IV

Superregnum: Eukaryota
Regnum: Plantae
Cladus: Angiosperms
Cladus: Eudicots
Cladus: Core eudicots
Cladus: Asterids
Cladus: Lamiids
Ordo: Lamiales

Familia: Plantaginaceae
Tribus: Gratioleae
Genera: Adenosma – Bacopa – Bythophyton – Cheilophyllum – Chodaphyton – Darcya – Deinostema – Dintera – Dizygostemon – Dopatrium – Encopella – Gratiola – Hydrotriche – Ildefonsia – Lapaea – Leucospora – Limnophila – Matourea – Mecardonia – Philcoxia – Schistophragma – Scoparia – Stemodia – Tetraulacium – Trapella – Umbraria

Name

Gratioleae Benth., Edwards's Bot. Reg. 21: 91. (1836).

Type genus: Gratiola L. (1753)

References
Primary references

Bentham, G. 1835. Edwards's Botanical Register 21: 91.

Additional references

Borsch, T., Müller, K. & Fischer, E. 2005. The Linderniaceae and Gratiolaceae are further lineages distinct from the Scrophulariaceae (Lamiales). Plant Biology (Stuttgart) 7(1): 1–2. ResearchGate
Colletta, G.D., Scatigna, A.V. & Souza, V.C. 2020. Assessment of Matourea pratensis (Plantaginaceae: Gratioleae) reveals an older name for Achetaria. Taxon 69(6): 1354-1360. DOI: 10.1002/tax.12293 Paywall ResearchGate Reference page.
Estes, D. & Small, R.L. 2008. Phylogenetic Relationships of the Monotypic Genus Amphianthus (Plantaginaceae Tribe Gratioleae) Inferred from Chloroplast DNA Sequences. Systematic Botany 33(1): 176–182. DOI: 10.1600/036364408783887375 JSTOR PDF Reference page.
Fischer, E., Schäferhoff, B. & Müller, K. 2013. The phylogeny of Linderniaceae – The new genus Linderniella, and new combinations within Bonnaya, Craterostigma, Lindernia, Micranthemum, Torenia and Vandellia. Willdenowia 43(2): 209–238. DOI: 10.3372/wi.43.43201 Open access Reference page.
Olmstead, R.G. (ed.) 2021. A Synoptical Classification of the Lamiales. Version 2.8.0 (in prog.). 20 pp. PDF Reference page. Accessed 14 Feb. 2014. As Gratiolieae.
Scatigna, A.V., Souza, V.C., Machado, R.M. & Simões, A.O. 2020. Lapaea (Plantaginaceae, Gratioleae), a new genus endemic to the Espinhaço Range (Brazil) with a remarkable red-flowered new species. Systematics and Biodiversity 18(8): 739-756. DOI: 10.1080/14772000.2020.1771470 Paywall ResearchGate Reference page.
Scatigna, A.V., Souza, V.C., Sosa, M.d.L.M., Colletta, G.D., Machado, R.M. & Simões, A.O. 2022. Phylogenetics of Gratioleae (Plantaginaceae): paraphyly of Stemodia and its implications for generic circumscriptions, with insights from floral evolution. Botanical Journal of the Linnean Society 200(2): 194–217. DOI: 10.1093/botlinnean/boac013 Paywall ResearchGate Reference page.

Links

International Plant Names Index. 2025. Gratioleae. Published online. Accessed: Jan. 26 2025. Reference page. pending

Vernacular names
العربية: غراتيولاوية