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Classification System: APG IV

Superregnum: Eukaryota
Regnum: Plantae
Cladus: Angiosperms
Cladus: Eudicots
Cladus: Core eudicots
Cladus: Asterids
Cladus: Lamiids
Ordo: Gentianales

Familia: Rubiaceae
Subfamilia: Cinchonoideae
Tribus: Guettardeae
Genus: AchilleanthusAntirheaArachnothryxBobeaChomeliaDichilantheGonzalaguniaGuettardaGuettardellaHodgkinsoniaMachaoniaMalaneaNeoblakeaNeolaugeriaOttoschmidtiaPittoniotisRogieraStenostomumTimoniusTinadendron


Guettardeae DC., Ann. Mus. Natl. Hist. Nat. 9: 217. (1807) as Guettardaceae.

Type genus: Guettarda L. Sp. Pl. 2: 991. (1753)


Candolle, A.P. de 1807. Annales du Muséum National d'Histoire Naturelle 9: 217.
Chavez, J.G., Meve, U., Nürk, N.M. & Liede-Schumann, S. 2021. Disentangling Antirhea (Rubiaceae): resurrection of Guettardella and description of the new genus Achilleanthus. Botanical Journal of the Linnean Society 197(1): 85-103. DOI: 10.1093/botlinnean/boab013 Paywall Reference page.
Manns, U. & Bremer, B. 2010. Towards a better understanding of intertribal relationships and stable tribal delimitations within Cinchonoideae s.s. (Rubiaceae). Molecular Phylogenetics and Evolution 56(1): 21-39. DOI: 10.1016/j.ympev.2010.04.002 PDF Reference page.
Rova, J.H.E., Delprete, P.G. & Bremer, B. 2009. The Rondeletia Complex (Rubiaceae): An Attempt to Use ITS, rps16, and trnL-F Sequence Data to Delimit Guettardeae, Rondeletieae, and Sections within Rondeletia. Annals of the Missouri Botanical Garden 96: 182-193. JSTOR PDF Reference page.
Guettardeae is a tribe of flowering plants in the family Rubiaceae and contains about 748 species in 14 genera. Its representatives are widespread geographically and are found in the tropics and subtropics.[1]

Currently accepted names[1][2][3][4]

Antirhea Comm. ex A.Juss. (38 sp)
Arachnothryx Planch. (107 sp)
Bobea Gaudich. (4 sp)
Chomelia Jacq. (79 sp)
Gonzalagunia Ruiz & Pav. (40 sp)
Guettarda L. (157 sp)
Hodgkinsonia F.Muell. (2 sp)
Machaonia Humb. & Bonpl. (31 sp)
Malanea Aubl. (40 sp)
Neoblakea Standl. (2 sp)
Rogiera Planch. (15 sp)
Stenostomum C.F.Gaertn. (49 sp)
Timonius Rumph. ex DC. (182 sp)
Tinadendron Achille (2 sp)


Abbottia F.Muell. = Timonius
Allenanthus Standl. = Machaonia
Anisomeris C.Presl = Chomelia
Bellermannia Klotzsch ex H.Karst. = Gonzalagunia
Bobaea A.Rich. = Bobea
Buena Cav. = Gonzalagunia
Bunophila Willd. ex Roem. & Schult. = Machaonia
Burneya Cham. & Schltdl. = Timonius
Cadamba Sonn. = Guettarda
Caruelina Kuntze = Chomelia
Caryococca Willd. ex Roem. & Schult. = Gonzalagunia
Cuatrecasasiodendron Steyerm. = Arachnothryx
Cunninghamia Schreb. = Malanea
Dicrobotryum Willd. ex Schult. = Guettarda
Donkelaaria Lem. = Guettarda
Duggena Vahl ex Standl. = Gonzalagunia
Edechia Loefl. = Guettarda
Erithalis G.Forst. = Timonius
Eupyrena Wight & Arn. = Timonius
Gonzalea Pers. = Gonzalagunia
Halesia P.Browne = Guettarda
Helospora Jacq. = Timonius
Javorkaea Borhidi & Jarai-Koml. = Arachnothryx
Laugeria L. = Guettarda
Laugeria Vahl ex Hook.f. = Stenostomum
Laugieria Jacq. = Guettarda
Matthiola L. = Guettarda
Microsplenium Hook.f. = Machaonia
Neolaugeria Nicolson = Stenostomum
Obbea Hook.f. = Bobea
Otocalyx Brandegee = Arachnothryx
Polyphragmon Desf. = Timonius
Porocarpus Gaertn. = Timonius
Resinanthus (Borhidi) Borhidi = Stenostomum
Rytidotus Hook.f. = Bobea
Sardinia Vell. = Guettarda
Schiedea A.Rich. = Machaonia
Siphonandra Turcz. = Arachnothryx
Sturmia C.F.Gaertn. = Stenostomum
Terebraria Sessé ex Kuntze = Stenostomum
Tertrea DC. = Machaonia
Tournefortiopsis Rusby = Guettarda
Viviania Raf. = Guettarda


"World Checklist of Rubiaceae". Retrieved May 2012. Check date values in: |accessdate= (help)
Bremer B (2009). "A review of molecular phylogenetic studies of Rubiaceae". Annals of the Missouri Botanical Garden. 96: 4–26. doi:10.3417/2006197.
Bremer B, Eriksson E (2009). "Time tree of Rubiaceae: phylogeny and dating the family, subfamilies, and tribes". International Journal of Plant Sciences. 170: 766–793. doi:10.1086/599077.
Robbrecht E, Manen J-F (2006). "The major evolutionary lineages of the coffee family (Rubiaceae, angiosperms). Combined analysis (nDNA and cpDNA) to infer the position of Coptosapelta and Luculia, and supertree construction based on rbcL, rps16, trnL-trnF and atpB-rbcL data. A new classification in two subfamilies, Cinchonoideae and Rubioideae". Systematic Geography of Plants. 76: 85–146.

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