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Classification System: APG IV

Superregnum: Eukaryota
Regnum: Plantae
Cladus: Angiosperms
Cladus: Eudicots
Cladus: Core eudicots
Cladus: Asterids
Cladus: Campanulids
Ordo: Apiales

Familia: Apiaceae
Subfamilia: Saniculoideae
Tribus: Saniculeae
Genus: Sanicula
Species: Sanicula mariversa

Sanicula mariversa Nagata & S.M.Gon

Systematic Botany; Quarterly Journal of the American Society of Plant Taxonomists 12:406. 1987
USDA, ARS, Germplasm Resources Information Network. Sanicula mariversa in the Germplasm Resources Information Network (GRIN), U.S. Department of Agriculture Agricultural Research Service. Accessed: 07-Oct-06.

Vernacular names
English: Waianae Range blacksnakeroot

Sanicula mariversa is a rare species of flowering plant in the family Apiaceae known by the common name Waianae Range black-snakeroot. It is endemic to Hawaii, where it is known only from the Waianae Mountains on the island of Oahu. It is threatened by the degradation of its habitat. It is a federally listed endangered species of the United States.

This plant produces erect, branching stems 40 to 70 centimeters long from a thick caudex. The leaves have leathery, dark green, heart-shaped or kidney-shaped blades up to 15 centimeters long by 23 wide which are palmately divided into three to five dissected lobes. The inflorescence is a cluster of many tiny yellow flowers, each flower measuring only about a 0.1 centimeter long. The fruit, measuring about half a centimeter long, is covered in hooked prickles.[2][3] This species is a monocarpic perennial, growing for several years, flowering and fruiting only once, then dying.[4] The plant withers after its growing season each year and remains dormant for the rest of the year, which is probably why it was not officially collected until 1985 nor described to science until 1987.[2] The plant was named for its residence on a ridge overlooking the ocean.[3]

This plant grows on the Makua-Keaau and Kaluaa-Lualualei Summit Ridges of the central Waianae Mountains.[4] The habitat is lower-elevation moist grassland.[1] The plants grow in deep soil and in cracks in steep rock cliffs. The surrounding ground has a layer of mosses and lichens. The area has been invaded by non-native species of grasses including soft chess (Bromus mollis) and annual fescue (Vulpia spp.). Another source of habitat degradation is the mountains' population of feral goats, which scale the slopes, breaking up the soil and causing erosion.[4]

As of 2006 there were 18 mature individuals and 170 immature plants remaining.[4]

Sanicula mariversa. The Nature Conservancy.
Nagata, K. M. and S. M. Gon. (1987). Sanicula mariversa (Apiaceae), a New Species from 'Ohikilolo Ridge, Wai'anae Mountains, O'ahu in the Hawaiian Archipelago. Systematic Botany 12(3) 406-09.
USFWS. Determination of Endangered Status for 26 Plants from the Waianae Mountains, Island of Oahu, Hawaii. Federal Register October 29, 1991.
USFWS. Sanicula mariversa Five-year Review. January 2008.

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