Cybocephalidae

Superregnum: Eukaryota
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Ordo: Coleoptera
Subordo: Polyphaga
Infraordo: Cucujiformia
Superfamilia: Cucujoidea

Familia: Cybocephalidae
Genera (8 + 1†): Cybocephalus – Endrodiellus – Hierronius – Horadion – Pastillodes – Pastillus – Pycnocephalus – Taxicephomerus – †Pastillocenicus
Name

Cybocephalinae Jacquelin du Val, 1858
References

Cline, A.R. et al. 2014: Molecular phylogeny of Nitidulidae: assessment of subfamilial and tribal classification and formalization of the family Cybocephalidae (Coleoptera: Cucujoidea). Systematic entomology, 39(4): 758–772. DOI: 10.1111/syen.12084 Reference page.

Additional references

Hisamatsu, S. 2013: A review of the Japanese Cybocephalidae (Coleoptera: Cucujoidea). Zootaxa 3616(3): 253–267. DOI: 10.11646/zootaxa.3616.3.3 Reference page.
Smith, T.R.; Cave, R.D. 2006: The Cybocephalidae (Coleoptera) of America north of Mexico. Annals of the Entomological Society of America 99(5): 776–792. DOI: 10.1603/0013-8746(2006)99[776:TCCOAN]2.0.CO;2 Paywall. Reference page.

Cybocephalidae is a family of sap, bark and fungus beetles in the order Coleoptera with a wide global distribution. The type genus Cybocephalus has more than 200 species in it and the entire family has about 220 species in all.[1][2][3][4] Many species are predators of armoured scale insects (Diaspididae).[5] There are four tarsal segments on all the legs. The body is only slightly longer than wide and very convex and shiny. They are small and about 1 to 3 mm long. The insect can roll into a ball like position with its downward facing head. The tarsomeres are lobed underneath. The family is sometimes treated as a subfamily within the Nitudulidae.[6]

The following genera have a 10 segmented antenna:

Apastillus Kirejtshuk & Mantic, 2015 monotypic, from Japan.
Theticephalus Kirejtshuk, 1988 from northwest Africa and Asia.
Pacicephalus Kirejtshuk & Mantic, 2015 from The Caroline Islands (northwest Pacific.)
Amedissia Kirejtshuk & Mantic, 2015, which is Neotropical.
Horadion Enrödy-Younga, 1976 from tropical Africa.

The following genera have an 11 segmented antenna:

Pycnocephalus Sharp, 1891 with 3 species from Central and South America.
Hierronius Enrödy-Younga, 1968 with 3 species endemic to Madeira and Canary Islands.
Pastillodes Enrödy-Younga, 1968 with 2 species from northwest Africa.
Taxicephomerus Kirejtshuk, 1994 monotypic, from Vietnam.
Pastillus Enrödy-Younga, 1962 with 2 Afrotropical species.
Endrodiellus Enrödy-Younga, 1962 which is monotypic from Madagascar.
Cybocephalus Erichson, 1844

References

"Cybocephalidae Family Information". BugGuide.net. Retrieved 2018-02-23.
"Cybocephalidae Report". Integrated Taxonomic Information System. Retrieved 2018-02-23.
"Cybocephalidae Overview". Encyclopedia of Life. Retrieved 2018-02-23.
"Browse Cybocephalidae". Catalogue of Life. Retrieved 2018-02-23.
Smith, Trevor Randall; Bailey, Rafique (2007). "A New Species of Cybocephalus (Coleoptera: Cybocephalidae) from Taiwan and a New Distribution Record for Cybocephalus nipponicus". The Coleopterists Bulletin. 61 (4): 503–508. JSTOR 27571054.

Lawrence, John F.; Slipinski, Adam (2013). Australian Beetles. Volume I. Morphology, Classification and Keys. CSIRO. pp. 325–327.

Further reading

Arnett, R.H. Jr.; Thomas, M. C.; Skelley, P. E.; Frank, J. H., eds. (2002). American Beetles, Volume II: Polyphaga: Scarabaeoidea through Curculionoidea. CRC Press.
Kellogg, Vernon L. (1905). American insects. H. Holt.
Arnett, Ross H. Jr. (2000). American Insects: A Handbook of the Insects of America North of Mexico. 2nd Edition. CRC Press. ISBN 0-8493-0212-9.
Leng, Charles W. (1920). Catalogue of the Coleoptera of America, North of Mexico. John D. Sherman, Jr.
Crotch, G.R. (1873). Check list of the Coleoptera of America, north of Mexico. Naturalists' Agency.
Capinera, John L., ed. (2008). Encyclopedia of Entomology. Springer. ISBN 978-1402062421.
Gillott, Cedric (1980). Entomology. Plenum Press. ISBN 0-306-40366-8.
Donald J. Borror; Roger Tory Peterson; Richard E. White (1998). A Field Guide to Insects. Houghton Mifflin.
Blatchley, W.S. (1910). An illustrated descriptive catalogue of the Coleoptera, beetles (exclusive of the Rhynchophora) known to occur in Indiana. Nature Pub.
Papp, Charles S. (1984). Introduction to North American Beetles. Entomography Pubns.
White, Richard E. (1983). Peterson Field Guides: Beetles. Houghton Mifflin.