Gnathaphanus

Superregnum: Eukaryota
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Ordo: Coleoptera
Subordo: Adephaga

Familia: Carabidae
Subfamilia: Harpalinae
Supertribus: Harpalitae
Tribus: Harpalini
Subtribus: Anisodactylina
Genus: Gnathaphanus

Species: G. aridus – G. chinensis – G. chujoi – G. froggatti – G. glamorgani – G. herbaceus – G. kansuensis – G. latus – G. licinoides – G. loeffleri – G. masudai – G. melbournensis – G. multipunctatus – G. parallelus – G. philippensis – G. picipes – G. porcatulus – G. pulcher – G. punctifer – G. rectangulus – G. riverinae – G. sexpunctatus – G. subolivaceus – G. vulneripennis – G. whitei
Name

Gnathaphanus Macleay, 1825

Type species: Gnathaphanus vulneripennis W.S. Macleay

Synonyms

Iwosiopelus Nakane & Ishida, 1959
Mirosarus Bates, 1878
Type species: Mirosarus insularis Bates, 1878 (monotypy)
Pachauchenius Macleay, 1864
Type species: Pachauchenius laeviceps W.J.MacLeay, 1864 (monotypy)

Gnathaphanus melbournensis

References
Primary references

Macleay, W.S. 1825. Annulosa Javanica, or an attempt to illustrate the natural affinities and analogies of the insects collected in Java by Thomas Horsfield, M. D. F. L. & G. S. and deposited by him in the museum of the honourable East-India Company. No. 1. Kingsbury, Parbury & Allen. London. DOI: 10.5962/bhl.title.65151 BHL Reference page. : 20
Bates, H.W. 1878. New genera and species of Carabidae from Tasmania. Cistula Entomologica 2: 317–326. BHL Reference page. : 319
Macleay, W.J. 1864. Descriptions of new genera and species of Coleoptera from Port Denison. Transactions of the Entomological Society of New South Wales 1: 106–130. BHL Reference page. : 116

Additional references

Darlington, P.J. 1968. The carabid beetles of New Guinea part III. Harpalinae (continued): Perigonini to Pseudomorphini. Bulletin of the Museum of Comparative Zoology 137(1): 1–253. BHL Reference page.
Larochelle, A. & Larivière, M.-C. 2005. Harpalini (Insecta: Coleoptera: Carabidae: Harpalinae). Fauna of New Zealand 53: 1–160. Reference page.

Links

Gnathaphanus – Taxon details on Biological Library (BioLib).
Gnathaphanus – Taxon details on Global Biodiversity Information Facility (GBIF).
ION
Nomenclator Zoologicus

Gnathaphanus is a genus of beetles in the family Carabidae, containing the following species:[1]

Gnathaphanus aridus Blackburn, 1892
Gnathaphanus froggatti (Macleay, 1888)
Gnathaphanus herbaceus Sloane, 1900
Gnathaphanus latus Sloane, 1900
Gnathaphanus licinoides Hope, 1842
Gnathaphanus melbournensis (Castelnau, 1867)
Gnathaphanus multipunctatus (Macleay, 1888)
Gnathaphanus philippensis (Chevrolat, 1841)
Gnathaphanus picipes (Macleay, 1864)
Gnathaphanus porcatulus (Macleay, 1888)
Gnathaphanus pulcher (Dejean, 1829)
Gnathaphanus punctifer (Castelnau, 1867)
Gnathaphanus riverinae Sloane, 1895
Gnathaphanus sexpunctatus (Macleay, 1888)
Gnathaphanus whitei Sloane, 1917

Parasites

In Australia, Gnathaphanus pulcher is parasitized by a species of mite, Eutarsopolipus pulcher Hajiqanbar & Seeman, 2021 which dwells under the elytra.[2]
References

"Gnathaphanus W.S. MacLeay, 1825: 20". Carabidae of the World. 2011. Retrieved 1 Jul 2011.
Katlav, Alihan; Hajiqanbar, Hamidreza; Riegler, Markus; Seeman, Owen D (2021). "Sheltered life beneath elytra: three new species of Eutarsopolipus (Acari, Heterostigmatina, Podapolipidae) parasitizing Australian ground beetles". Parasite. EDP Sciences. 28: 75. doi:10.1051/parasite/2021069. ISSN 1776-1042. open access