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Superregnum: Eukaryota
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Superphylum: Deuterostomia
Phylum: Chordata
Cladus: Craniata
Subphylum: Vertebrata
Infraphylum: Gnathostomata
Superclassis: Tetrapoda
Classis: Amphibia
Subclassis: Lissamphibia
Ordo: Anura

Familia: Microhylidae
Subfamilia: Cophylinae
Genus: Cophyla
Species (6): C. alticola – C. ando – C. barbouri – C. berara – C. cowanii – C. fortuna – C. grandis – C. karenae – C. maharipeo – C. mavomavo – C. milloti – C. noromalalae – C. occultans – C. olgae – C. occultans – C. phyllodactyla – C. pollicaris – C. puellarum – C. rava – C. tetra – C. tsaratananaensis – C. tuberifera
Name

Cophyla Boettger, 1880: 281

Type species: Cophyla phyllodactyla Boettger, 1880, by monotypy.
References
Primary references

Boettger, O. 1880. Diagnoses reptilium et batrachiorum novorum a Carolo Ebenau in insula Nossi-Bé Madagascariensis lectorum. Zoologischer Anzeiger 3: 279–283. BHL

Additional references

Vences, M., Andreone, F. & Glaw, F. 2005. A new microhylid frog of the genus Cophyla from a transitional forest in northwestern Madagascar. African Zoology 40(1): 143–149. Reference page.
Rakotoarison, A., Crottini, A., Müller, J., Rödel, M-O., Glaw, F. & Vences, M. 2015. Revision and phylogeny of narrow-mouthed treefrogs (Cophyla) from northern Madagascar: integration of molecular, osteological, and bioacoustic data reveals three new species. Zootaxa 3937(1): 61–89. DOI: 10.11646/zootaxa.3937.1.3. Reference page.

Links

Amphibian Species of the World 5.1 Cophyla access date 16 June 2008

Vernacular names
English: Whistling Treefrogs

Cophyla is a genus of microhylid frogs endemic to Madagascar.[1][2]
Species

The following species are recognised in the genus Cophyla:[1]

Cophyla alticola (Guibé, 1974)
Cophyla ando (Scherz, Köhler, Vences, and Glaw, 2019)
Cophyla barbouri (Noble, 1940)
Cophyla berara Vences, Andreone, and Glaw, 2005
Cophyla cowanii (Boulenger, 1882)
Cophyla fortuna Rakotoarison, Scherz, Bletz, Razafindraibe, Glaw, and Vences, 2019
Cophyla grandis (Boulenger, 1889)
Cophyla karenae (Rosa, Crottini, Noel, Rabibisoa, Raxworthy, and Andreone, 2014)
Cophyla laetus (Rakotoarison, Scherz, Köhler, Ratsoavina, Hawlitschek, Megson, Vences & Glaw, 2020)
Cophyla maharipeo Rakotoarison, Crottini, Müller, Rödel, Glaw, and Vences, 2015
Cophyla mavomavo (Andreone, Fenolio, and Walvoord, 2003)
Cophyla milloti (Guibé, 1950)
Cophyla noromalalae Rakotoarison, Crottini, Müller, Rödel, Glaw, and Vences, 2015
Cophyla occultans (Glaw and Vences, 1992)
Cophyla olgae (Rakotoarison, Glaw, Vieites, Raminosoa, and Vences, 2012)
Cophyla phyllodactyla Boettger, 1880
Cophyla pollicaris (Boulenger, 1888)
Cophyla puellarum Rakotoarison, Crottini, Müller, Rödel, Glaw, and Vences, 2015
Cophyla rava (Glaw, Köhler, and Vences, 2012)
Cophyla tetra (Andreone, Fenolio, and Walvoord, 2003)
Cophyla tsaratananaensis Guibé, 1974
Cophyla tuberifera (Methuen, 1920)

References

Frost, Darrel R. (2015). "Cophyla Boettger, 1880". Amphibian Species of the World: an Online Reference. Version 6.0. American Museum of Natural History. Retrieved 9 June 2015.
Rakotoarison, Andolalao; Crottini, Angelica; Müller, Johannes; Rödel, Mark-Oliver; Glaw, Frank; Vences, Miguel (2015). "Revision and phylogeny of narrow-mouthed treefrogs (Cophyla) from northern Madagascar: integration of molecular, osteological, and bioacoustic data reveals three new species". Zootaxa. 3937 (1): 61–89. doi:10.11646/zootaxa.3937.1.3. PMID 25947461.

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