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Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Superphylum: Deuterostomia
Phylum: Chordata
Subphylum: Vertebrata
Infraphylum: Gnathostomata
Megaclassis: Osteichthyes
Cladus: Sarcopterygii
Cladus: Rhipidistia
Cladus: Tetrapodomorpha
Cladus: Eotetrapodiformes
Cladus: Elpistostegalia
Superclassis: Tetrapoda
Cladus: Reptiliomorpha
Cladus: Amniota
Classis: Reptilia
Cladus: Eureptilia
Cladus: Romeriida
Subclassis: Diapsida
Cladus: Sauria
Cladus: Archelosauria
Division: Pan-Testudines
Division: Testudinata
Ordo: Testudines
Subordo: Cryptodira
Superfamilia: Testudinoidea

Familia: Testudinidae
Genus: Manouria
Species: M. emys - M. impressa - †M. oyamai
Name

Manouria Gray 1852
Vernacular names
日本語: ムツアシガメ属
ไทย: เต่าหก, เต่าเดือย

Manouria is a genus of tortoises in the family Testudinidae. The genus was erected by John Edward Gray in 1854.
Species

The following five species are recognized as being valid, two of which are extant,[1] and three of which are extinct:

Manouria emys (Schlegel & S. Müller, 1844) – Asian forest tortoise
Manouria impressa (Günther, 1882) – impressed tortoise
†Manouria sondaari Karl & Staesche, 2007 – a giant land tortoise from Luzon Island, Philippines[2][3] however, Rhodin et al. (2015) transferred this species to the genus Megalochelys.[4]
†Manouria punjabiensis (Lydekker, 1889) – a fossil tortoise from Siwaliks, India[4]
†Manouria oyamai Takahashi, Otsuka & Hirayama , 2003 – a fossil tortoise from Ryukyu Islands, Japan[4]

Nota bene: A binomial authority in parentheses indicates that the species was originally described in a genus other than Manouria.
References

"Manouria". The Reptile Database. www.reptile-database.org.
Karl, Hans-Volker; Staesche, Ulrich (2007). "Fossile Riesen-Landschildkröten von den Philippinen und ihre paläogeographische Bedeutung [= Fossil Giant Land Tortoises from the Philippines and their paleogeographic importance]". Geologischesahrbuch A 160: 171–197. (Manouria sondaari, new species). (in German with abstract in English).
Staesche, Ulrich (coordinator) (2007). Fossile Schildkröten aus vier Ländern in drei Kontinenten: Deutschland, Türkei, Niger, Philippen [= Fossil Turtles from Four Countries on Three Continents: Germany, Turkey, Niger, and the Philippines]. Geologisches Jahrbuch, Reihe B, Heft 98 [= Series B, Issue 98]. 197 pp. ISBN 978-3-510-95967-9. http://www.schweizerbart.de/publications/detail/artno/186029800 (in German).

Anders G.J. Rhodin; Scott Thomson; Georgios L. Georgalis; Hans-Volker Karl; Igor G. Danilov; Akio Takahashi; Marcelo S. de la Fuente; Jason R. Bourque; Massimo Delfino; Roger Bour; John B. Iverson; H. Bradley Shaffer; Peter Paul van Dijk (2015). "Turtles and Tortoises of the World During the Rise and Global Spread of Humanity: First Checklist and Review of Extinct Pleistocene and Holocene Chelonians" (PDF). Chelonian Research Monographs. 5 (8): 000e.1–66. doi:10.3854/crm.5.000e.fossil.checklist.v1.2015.

Further reading

Gray, J. E. (1854). "Description of a New Genus and some New Species of Tortoises". Proceedings of the Zoological Society of London. 1852: 133–135. (Manouria, new genus, p. 133).

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