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Superregnum: Eukaryota
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: †Trilobitomorpha
Classis: Trilobita
Ordo: Agnostida
Subordo: Agnostina
Superfamilia: Agnostoidea

Familia: Agnostidae
Subfamilia: Agnostinae
Genus: Acmarhachis

Species: †A. acuta – †A. apicula – †A. elegans – †A. karatauensis – †A. kindlei – †A. longispinus – †A. punctatus – †A. typicalis – †A. whittingtoni

Acmarhachis Resser, 1938


Cyclagnostus Lermontova, 1940


Resser C.E., 1938. Spec. Papers Geol. Soc. Amer., 15, page 47.


Acmarhachis – Taxon details on Fossilworks.
Acmarhachis – Taxon details on Global Biodiversity Information Facility (GBIF).
Acmarhachis – Taxon details on Interim Register of Marine and Non-marine Genera (IRMNG).

Acmarhachis is a genus of trilobite in the order Agnostida, which lived in what are now Australia (Queensland, Tasmania), Canada (British Columbia, Newfoundland, Northwest Territories), China (Anhui), Kazakhstan, Russia (Kharaulakh), and the US (Alaska, Alabama, Nevada, Maryland, Vermont). It was described by Resser in 1938, and the type species is Acmarhachis typicalis.[3]

A. typicalis occurs in the Cambrian of Kazakhstan (Kyrshabakty River, Lejopyge laevigata-trilobite zone, Zhumabai Formation, Paibian and Cambrian Stage 9, 43.5° N, 70.0° E).[4]
A. whittingtoni is known from the Upper Cambrian of Canada (Hickey Cove, Dunderbergia-trilobite zone, Downes Point Member, Shallow Bay Formation, Paibian, Newfoundland, 49.8° N, 57.9° W).[5]


Like all Agnostida, Acmarhachis is diminutive and the headshield (or cephalon) and tailshield (or pygidium) are of approximately the same size (or isopygous) and outline. As in all Agnostina, Acmarhachis has two thorax segments. When Agnostina are enrolled, a tiny opening between the cephalon and the thorax becomes visible (the cephalothoracic aperture) that uniquely distinguishes this group from the Eodiscina. A feature shared with all Agnostoidea is that the frontal lobe of the central raised area of the cephalon (or glabella) is not laterally expanded compared to the backward lobe, like in the Condylopygoidea. In Acmarhachis the furrow at midline between the front of the glabella and the border is weak or absent. The glabella has a long ogival to subquadrate front lobe, and the furrow defining it at its back is straight. The back lobe of the glabella is accompanied by two triangular basal lobes of moderate to large size. On the pygidium the axis or rhachis reaches the border. It is divided along its length in three lobes. It is shaped a bit like an amphora, the back pointed and strongly constricted across the middle lobe. The middle lobe also carries a node where it contacts the posterior lobe.[2]


Arthropoda - Klass trilobity (Class Trilobita). EV Lermontova, Atlas of the Leading Forms of the Fossil Faunas of the USSR, 1940
Whittington, H. B. et al. Part O, Treatise on Invertebrate Paleontology. Revised, Volume 1 – Trilobita – Introduction, Order Agnostida, Order Redlichiida. 1997
Available Generic Names for Trilobites P.A. Jell and J.M. Adrain.
Ergaliev, G.K.; Ergaliev, F.G. (2008). Agnostidy srednego i verkhnego Kembriya Aksayskogo Gosudarstvennogo Geologicheskogo Zakaznika v yushnom Kazakhstane (Kyrshabakty, Malyy Karatau) [Middle and Upper Cambrian Agnostida from the Aksai National Geological Reserve in southern Kazakhstan (Kyrshabakty River, Malyy Karatau Range)]. pp. 1–359. cited in Paleobiological Database. "Acmarhachis whittingtoni". Retrieved 25 October 2013.
Westrop, S.R.; Eoff, .J.D. (2012). "Late Cambrian (Furongian; Paibian, Steptoean) agnostoid arthropods from the Cow Head Group, western Newfoundland". Journal of Paleontology. 86 (2): 201–237. doi:10.1666/11-034.1. cited in Paleobiological Database. "Acmarhachis whittingtoni". Retrieved 25 October 2013.


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