Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: ParaHoxozoa
Cladus: Bilateria
Cladus: Nephrozoa
Superphylum: Deuterostomia
Phylum: Chordata
Cladus: Olfactores
Subphylum: Vertebrata
Infraphylum: Gnathostomata
Megaclassis: Osteichthyes
Cladus: Sarcopterygii
Cladus: Rhipidistia
Cladus: Tetrapodomorpha
Cladus: Eotetrapodiformes
Cladus: Elpistostegalia
Superclassis: Tetrapoda
Cladus: Reptiliomorpha
Cladus: Amniota
Cladus: Sauropsida
Cladus: Reptilia
Cladus: Eureptilia
Cladus: Romeriida
Subclassis: Diapsida
Cladus: Neodiapsida
Cladus: Sauria
Cladus: Archelosauria
Cladus: Archosauromorpha
Cladus: Crocopoda
Cladus: Archosauriformes
Cladus: Eucrocopoda
Cladus: Archosauria
Cladus: Avemetatarsalia
Cladus: Ornithodira
Cladus: Dinosauromorpha
Cladus: Dinosauriformes
Cladus: Dracohors
Cladus: Dinosauria
Cladus: Saurischia
Cladus: Theropoda
Cladus: Neotheropoda
Cladus: Averostra
Cladus: Tetanurae
Cladus: Avetheropoda
Cladus: Coelurosauria
Cladus: Tyrannoraptora
Cladus: Maniraptoromorpha
Cladus: Maniraptoriformes
Cladus: Maniraptora
Cladus: Pennaraptora
Cladus: Paraves
Cladus: Eumaniraptora
Cladus: Avialae
Subclassis: Aves
Ordo: Passeriformes
Subordo: Passeri
Infraordo: Passerida
Superfamilia: Sylvioidea
Familia: Pycnonotidae
Genus: Hemixos
Species: H. castanonotus - H. flavala
Name
Hemixos Blyth, 1845
References
The Journal of the Asiatic Society of Bengal 14 p. 572
Hemixos is a songbird genus in the bulbul family, Pycnonotidae.[1]
Taxonomy and systematics
Established by Edward Blyth in 1845 for the newly discovered ashy bulbul (H. flavala),[2] this genus contains four extant species.[1]
Some treatments merge the genus into Hypsipetes, often together with the rest of the traditional "Hypsipetes group" of bulbuls: Iole, Ixos, Microscelis and Tricholestes. But in this case, the closely related genera Alophoixus and Setornis would probably also have to be included, and as soon as the earliest described genus, Ixos, is merged with another its name would apply.[3]
In fact, Hemixos is not particularly close to Hypsipetes, and a merger is not well justified. mtDNA NADH dehydrogenase subunits 2 and 3 and nDNA β-fibrinogen intron 7 sequence data puts it closer to (but still well distant from) the streaked bulbul (Ixos malaccensis). But whether that species represents the core group of Ixos – around its type species I. virescens (Sunda bulbul or green-winged bulbul) –, or a distinct lineage worthy of separation in a new genus – in which case Hypsipetes might be merged into the core group of Ixos – has not been studied. In any case, though minor, the Hemixos lineage with its stark white throat and light wing patches seems well distinct.[4]
Extant species
Four species are currently recognised:[1]
Cream-striped Bulbul (Hemixos leucogrammicus; moved from Pycnonotus following molecular phylogenetic analyses)
Ashy bulbul (Hemixos flavala)
Cinereous bulbul (Hemixos cinereus)
Chestnut bulbul (Hemixos castanonotus)
Former species
Previously, some authorities also classified the following species (or subspecies) as species within the genus Hemixos:
Sumatran bulbul (as Hemixus sumatranus)[5]
Footnotes
Gill, F.; Donsker, D.; Rasmussen, P. (eds.). "Family Pycnonotidae". IOC World Bird List. Version 10.2. International Ornithological Congress. Retrieved 5 August 2020.
Gregory (2000)
Gregory (2000), Pasquet et al. (2001), Moyle & Marks (2006)
Pasquet et al. (2001), Moyle & Marks (2006)
"Ixos virescens sumatranus (Sumatran Bulbul) - Avibase". avibase.bsc-eoc.org. Retrieved 2017-09-23.
References
Media related to Hemixos at Wikimedia Commons
Gregory, Steven M. (2000): Nomenclature of the Hypsipetes Bulbuls (Pycnonotidae). Forktail 16: 164–166. PDF fulltext
Moyle, Robert G. & Marks, Ben D. (2006): Phylogenetic relationships of the bulbuls (Aves: Pycnonotidae) based on mitochondrial and nuclear DNA sequence data. Mol. Phylogenet. Evol. 40(3): 687–695. doi:10.1016/j.ympev.2006.04.015 (HTML abstract)
Pasquet, Éric; Han, Lian-Xian; Khobkhet, Obhas & Cibois, Alice (2001): Towards a molecular systematics of the genus Criniger, and a preliminary phylogeny of the bulbuls (Aves, Passeriformes, Pycnonotidae). Zoosystema 23(4): 857–863. PDF fulltext
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