Superregnum: Eukaryota
Cladus: Orthokaryota
Cladus: Neokaryota
Cladus: Scotokaryota
Cladus: Podiata
Cladus: Amorphea
Cladus: Obazoa
Cladus: Opisthokonta
Cladus: Holozoa
Cladus: Filozoa
Cladus: Choanozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: ParaHoxozoa
Cladus: Bilateria
Cladus: Nephrozoa
Superphylum: Deuterostomia
Phylum: Chordata
Cladus: Olfactores
Subphylum: Vertebrata
Infraphylum: Gnathostomata
Cladus: Eugnathostomata
Cladus: Osteichthyes
Cladus: Sarcopterygii
Cladus: Rhipidistia
Cladus: Tetrapodomorpha
Cladus: Eotetrapodiformes
Cladus: Elpistostegalia
Cladus: Tetrapoda
Cladus: Reptiliomorpha
Cladus: Amniota
Cladus: Sauropsida
Cladus: Reptilia
Cladus: Eureptilia
Cladus: Romeriida
Subclassis: Diapsida
Cladus: Neodiapsida
Cladus: Sauria
Cladus: Archelosauria
Cladus: Archosauromorpha
Cladus: Crocopoda
Cladus: Archosauriformes
Cladus: Eucrocopoda
Cladus: Archosauria
Cladus: Avemetatarsalia
Cladus: Ornithodira
Cladus: Dinosauromorpha
Cladus: Dinosauriformes
Cladus: Dracohors
Cladus: Dinosauria
Cladus: Saurischia
Cladus: Theropoda
Cladus: Neotheropoda
Cladus: Averostra
Cladus: Tetanurae
Cladus: Avetheropoda
Cladus: Coelurosauria
Cladus: Tyrannoraptora
Cladus: Maniraptoromorpha
Cladus: Maniraptoriformes
Cladus: Maniraptora
Cladus: Pennaraptora
Cladus: Paraves
Cladus: Eumaniraptora
Cladus: Avialae
Subclassis: Aves
Ordo: Passeriformes
Subordo: Passeri
Infraordo: Passerida
Superfamilia: Sylvioidea
Familia: Leiothrichidae
Genera: Actinodura – Babax – Crocias – Cutia – Garrulax – Heterophasia – Kupeornis – Leiothrix – Liocichla – Minla – Phyllanthus – Pterorhinus – Trochalopteron – Turdoides –
Name
Leiothrichidae Swainson, 1832
Typus
Leiothrix
Swainson, 1832
References
Fauna boreali-americana 2: 232, 490, pro subfamilia, "Leiotrichanæ".
IOC World Bird List 7.1
Vernacular names
Deutsch: Häherlinge
English: Laughingthrushes
中文: 噪鹛科
The laughingthrushes are a family, Leiothrichidae, of Old World passerine birds. They are diverse in size and coloration. These are birds of tropical areas, with the greatest variety in Southeast Asia and the Indian subcontinent. The entire family used to be included in the Old World babbler family Timaliidae.
Characteristics
They are small to medium-sized birds. They have strong legs, and many are quite terrestrial. They typically have generalised bills, similar to those of a thrush. Most have predominantly brown plumage, with minimal difference between the sexes, but many more brightly coloured species also exist.[2]
This group is not strongly migratory, and most species have short rounded wings, and a weak flight. They live in lightly wooded or scrubland environments, ranging from swamp to near-desert. They are primarily insectivorous, although many will also take berries, and the larger species will even eat small lizards and other vertebrates.[2]
Taxonomy
The family Leiothrichidae was introduced (as a subfamily Leiotrichanae) by the English naturalist William Swainson in 1832.[3] A comprehensive molecular phylogenetic study of the family published in 2018 led to substantial revision of the taxonomic classification.[1] The laughingthrushes in the genus Garrulax were found to belong to three separate clades that had diverged in the Miocene 7-9 million year ago. The genus was therefore split with Garrulax restricted to one clade and the genera Pterorhinus and Ianthocincla resurrected for the other two clades. The genus Turdoides was also split and species moved into the resurrected genus Argya.[1][4]
In a separate change, the crocias were moved to the genus Laniellus Swainson, 1832 which has priority over Crocias Temminck, 1836.[4][5][6]
List of genera
The family contains 133 species in 16 genera:[4]
Grammatoptila – striated laughingthrush
Cutia – cutias (2 species)
Laniellus – crocias (2 species)
Trochalopteron – laughingthrushes (19 species)
Actinodura – barwings and minlas (9 species)
Montecincla – laughingthrushes (4 species)
Minla – red-tailed minla
Leioptila – rufous-backed sibia
Leiothrix – (2 species)
Liocichla - liocichlas (5 species)
Heterophasia - sibias (7 species)
Argya – mainly babblers (16 species) – previous placed in Turdoides
Turdoides – babblers (19 species)
Garrulax – laughingthrushes (14 species)
Ianthocincla – laughingthrushes (8 species) – previously placed in Garrulax
Pterorhinus – laughingthrushes and babaxes (23 species) – previously placed in Garrulax
References
Cibois, A.; Gelang, M.; Alström, P.; Pasquet, E.; Fjeldså, J.; Ericson, P.G.P.; Olsson, U. (2018). "Comprehensive phylogeny of the laughingthrushes and allies (Aves, Leiothrichidae) and a proposal for a revised taxonomy". Zoologica Scripta. 47 (4): 428–440. doi:10.1111/zsc.12296.
Perrins, C. (1991). Forshaw, Joseph (ed.). Encyclopaedia of Animals: Birds. London: Merehurst Press. pp. 188–190. ISBN 1-85391-186-0.
Swainson, William John; Richardson, J. (1831). Fauna boreali-americana, or, The zoology of the northern parts of British America. Part 2. The Birds. London: J. Murray. p. 490. The title page bears the year 1831 but the volume did not appear until 1832.
Gill, Frank; Donsker, David; Rasmussen, Pamela, eds. (January 2021). "Laughingthrushes and allies". IOC World Bird List Version 11.1. International Ornithologists' Union. Retrieved 18 June 2021.
Gregory, S.M.S.; Dickinson, E. (2012). "An assessment of three little‐noticed papers on avian nomenclature by G.N. Kashin during 1978‐1982". Zootaxa. 3340: 44-58 [51].
Dickinson, E.C.; Christidis, L., eds. (2014). The Howard & Moore Complete Checklist of the Birds of the World. Volume 2: Passerines (4th ed.). Eastbourne, UK: Aves Press. p. 548. ISBN 978-0-9568611-2-2.
Further reading
Cibois, A. (2003). "Mitochondrial DNA phylogeny of babblers (Timaliidae)". Auk. 120: 35–54. doi:10.1642/0004-8038(2003)120[0035:MDPOBT]2.0.CO;2.
Collar, N. J., and C. Robson. 2007. Family Timaliidae (babblers). Pages 70–291 in J. del Hoyo, A. Elliott, and D.A. Christie (editors), Handbook of the Birds of the World Volume 12: Picathartes to Tits and Chickadees. Lynx Edicions, Barcelona.
Gelang, M., A. Cibois, E. Pasquet, U. Olsson, P. Alström and P.G.P. Ericson. 2009. Phylogeny of babblers (Aves, Passeriformes): major lineages, family limits and classification. Zoologica Scripta 38: 225–236.
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