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Tetraogallus himalayensis

Tetraogallus himalayensis, Photo: Michael Lahanas

Superregnum: Eukaryota
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Superphylum: Deuterostomia
Phylum: Chordata
Subphylum: Vertebrata
Infraphylum: Gnathostomata
Megaclassis: Osteichthyes
Superclassis: Sarcopterygii
Superclassis: Tetrapoda
Cladus: Reptiliomorpha
Cladus: Amniota
Classis: Reptilia
Cladus: Eureptilia
Cladus: Romeriida
Subclassis: Diapsida
Cladus: Sauria
Infraclassis: Archosauromorpha
Cladus: Crurotarsi
Divisio: Archosauria
Cladus: Avemetatarsalia
Cladus: Ornithodira
Subtaxon: Dinosauromorpha
Cladus: Dinosauriformes
Cladus: Dracohors
Cladus: Dinosauria
Ordo: Saurischia
Cladus: Eusaurischia
Cladus: Theropoda
Cladus: Neotheropoda
Cladus: Averostra
Cladus: Tetanurae
Cladus: Avetheropoda
Cladus: Coelurosauria
Cladus: Tyrannoraptora
Cladus: Maniraptoromorpha
Cladus: Maniraptoriformes
Cladus: Maniraptora
Cladus: Pennaraptora
Cladus: Paraves
Cladus: Eumaniraptora
Cladus: Avialae
Infraclassis: Aves
Cladus: Euavialae
Cladus: Avebrevicauda
Cladus: Pygostylia
Cladus: Ornithothoraces
Cladus: Euornithes
Cladus: Ornithuromorpha
Cladus: Ornithurae
Cladus: Carinatae
Parvclassis: Neornithes
Cohors: Neognathae
Cladus: Pangalloanserae
Cladus: Galloanseres
Ordo: Galliformes

Familia: Phasianidae
Subfamilia: Perdicinae
Genus: Tetraogallus
Species: Tetraogallus himalayensis
Subspecies: T. h. bendi - T. h. grombczewskii - T. h. himalayensis - T. h. incognitus - T. h. koslowi - T. h. saurensis - T. h. sewerzowi

Tetraogallus himalayensis G.R. Gray, 1843

Proceedings of the Zoological Society of London (1842) Pt10 no.114 p. 105
Vernacular names
English: Himalayan Snowcock
español: Perdigallo himalayo
日本語: ヒマラヤセッケイ

The Himalayan snowcock (Tetraogallus himalayensis) is a snowcock in the pheasant family Phasianidae found across the Himalayan ranges and parts of the adjoining Pamir range of Asia. It is found on alpine pastures and on steep rocky cliffs where they will dive down the hill slopes to escape. It overlaps with the slightly smaller Tibetan snowcock in parts of its wide range. The populations from different areas show variations in the colouration and about five subspecies have been designated. They were introduced in the mountains of Nevada in the United States in the 1960s and a wild population has established in the Ruby Mountains.

Illustration by J. G. Keulemans (1891)

The Himalayan snowcock is a large grey partridge-like bird, 55–74 cm (22–29 in) in length and weighing 2–3.1 kg (4.4–6.8 lb).[2][3] The head pattern has a resemblance to that of the smaller and well marked chukar partridge. The white throat and sides of the head are bordered by chestnut moustachial stripe and a dark broad chestnut band stretching from the eye over the ear, expanding into the collar. The upper parts are grey, with feathers of the rump and the wings are bordered with rufous. The upper breast is grey with dark crescent bars. The lower breast plumage is dark grey, and the sides of the body are streaked with black, chestnut and white. The undertail coverts are white. The legs and orbital skin are yellow. Sexes are alike in plumage, but the female is smaller and lacks the large tarsal spur of the male. In flight, from above, the white primaries tipped in black and the rufous outer tail feathers make it distinctive. The Tibetan snowcock has a wing pattern with white trailing edge to the secondaries that contrasts with the grey wings.[4][5][6][7][8][9]
Taxonomy and systematics

Around 1841 specimen of a bird was brought to the menagerie of the Zoological Society of London, presented by E. W. Bonham, consul at Tabrez. George Robert Gray noted that it matched a bird illustrated (plate 76) in Jardine and Selby's "Illustrations of Ornithology" which he noted should be called as Tetraogallus caucasicus and another illustration plate 141 which was said to be the male of the one in plate 76 was separated based on the geography and given the name of Tetraogallus himalayensis.[10] The main populations are separated by the Taklamakan desert and the separation has been estimated to have happened after the Pleistocene glaciations (1 mya).[11] The large distribution range is fragmented and these disjunct populations show variations in plumage that have been named as subspecies. Although several subspecies have been described, not all are recognized.[12] A subspecies described as sauricus was the subject of considerable nomenclatural debate.[13][14]

The nominate subspecies was described by G. R. Gray in 1843 and refers to the populations from eastern Afghanistan extending to Ladakh and the central Himalayas of Nepal.
incognitus described by Zarudny in 1911 from the mountains of southern Tadjikistan and northern Afghanistan, is overall much paler with very light chestnut and black markings on underside and buffier.[5](bendi described by Walter Norman Koelz is usually included in this[5][15][16])
sewerzowi Zarudny, 1910 – Tien Shan to Zaysan (Kazakhstan) east to Xinjiang (not recognized by Ernst Hartert[16])
grombczewskii Bianchi, 1898 – Kunlun Mountains
koslowi Bianchi, 1898 – Nan Shan and Ching Hai Ku Mountains of Qinghai and S Gansu

Distribution and status

Himalayan snowcock frequents alpine pastures and steep ridges of mountains of Central and South Asia above the treeline and near the snowline. In the Himalayas, it is found between 4000 and 5000 m elevation in summer, descending to 2400 m during severe winters.[17] Since the Himalayan snowcock has a large distribution range and no visible declines in population, it has been considered a species of "least concern" by the IUCN.[1]

In 1961 the similarity of the Himalayan landscape to the Nevada region was noted and the Himalayan snowcock was considered as a good game bird for introduction by the Nevada Fish and Game Commission. The Commission then approached the President of Pakistan for some birds. These were wild trapped in Hunza and early shipments faced heavy losses after which birds were locally reared at Mason Valley game farm and over a 15-year period (1965–1979) more than 2000 birds were released into the wild. A wild population more than 200 to 500 birds has established itself in the Ruby Mountains, where they forage above the treeline.[18]
Behaviour and ecology
Egg, Collection Museum Wiesbaden

Himalayan snowcocks are gregarious when not breeding, moving around in small groups. Several groups may inhabit the same hill. They keep entirely to open country and seem to prefer rocky hill-sides. They feed on grass, shoots, berries and seeds.[19] In the mornings the birds fly downhill to drink water. When approached from below their level, they attempt to climb up the slopes on foot and when approached from above they dive down the valleys on open wings. In India, the breeding season is in summer, April to June. It is silent in winter but in spring, its call is a familiar part of the landscape. The song is a loud whistle with three parts with the tone ascending. They also make a rising, shrill piping call.[5]

When feeding they walk slowly up hill, picking up the tender blades of grass and young shoots of plants on the way. They have been noted feeding on the berries of Ephedra, leaves of Artemisia, grass shoots, bulbs and the heads of a rye-like grass.[17] In the Hunza range, they have been observed to prefer Sibbaldia cuneata.[20] Once they reach the top of a ridge of the hill, they fly off to adjacent hill, alighting some distance down, and again picking their way upwards. When walking, they cock their tails showing the white under tail coverts. They are generally wary and when disturbed run uphill and then launch themselves from the crests in flight, getting up considerable speed.[21] They are more prone to predation on the alpine pastures than on steep slopes and flocking helps them to keep more eyes out for predators allowing them to forage more efficiently.[22] In the Hunza range, flock sizes tended to be larger in rocky habitats, where they risked being attacked by Golden Eagles, than on grassy meadows.[20]

The breeding season is summer, April to June. During courtship, the male crouches low down to the ground with wings slightly spread, tail depressed and feathers slightly ruffled. Then, he runs backwards and forwards in front of the hen or in circles. The nest is a bare ground scrape sheltered under a stone or bush, preferably close to the crest of a ridge on the leeward side. About 5 to 12 long oval eggs are laid which are a stony olive or brown colour and spotted throughout with red or brown.[19] The eggs are incubated only by the female. The male is monogamous, staying in the vicinity of the nest often seen perched on some elevated rock and keeping a watch against intruders. When disturbed, the cock warns the female with a loud whistle. If caught unaware in the nest, the hen bird will not leave the nest until approached very close. The eggs hatched in an incubator after about 27–28 days.[17][23][24]

Adults are sometimes preyed on by golden eagles.[6][25] Several species of endoparasitic Acanthocephala and Nematoda such as Hispaniolepis fedtschenkoi, have been described from the species.[26][27]

BirdLife International (2018). "Tetraogallus himalayensis". IUCN Red List of Threatened Species. 2018: e.T22678673A132049858. doi:10.2305/IUCN.UK.2018-2.RLTS.T22678673A132049858.en. Retrieved 12 November 2021.
CRC Handbook of Avian Body Masses by John B. Dunning Jr. (Editor). CRC Press (1992), ISBN 978-0-8493-4258-5.
[1] Archived 25 April 2012 at the Wayback Machine (2011).
Oates, EW (1898). A manual of the Game birds of India. Part 1. A J Combridge, Bombay. pp. 201–204.
Rasmussen PC; JC Anderton (2005). Birds of South Asia: The Ripley Guide. Volume 2. Smithsonian Institution & Lynx Edicions. p. 119.
Jerdon, TC (1864). The birds of India. Volume 3. George Wyman and Co, Calcutta. pp. 549–554.
Blanford, WT (1898). The Fauna of British India, Including Ceylon and Burma. Birds. Volume 4. Taylor and Francis, London. pp. 143–144.
Baker, ECS (1928). The Fauna of British India, Including Ceylon and Burma. Birds. Volume 5 (2nd ed.). Taylor and Francis, London. pp. 426–428.
Ogilvie-Grant, WR (1896). A hand-book to the Game-birds. Volume 1. Edward Lloyd, London. pp. 86–89.
Gray GR (1843). "[Letter to curator]". Proceedings of the Zoological Society of London (Part 10): 105.
Ruan Luzhang; An Bei; Niclas Backstrom; Luo Huaxing; Wen Longying; Zhang Lixun; Liu Naifa (2010). "Phylogeographic structure and gene flow of Himalayan snowcock (Tetraogallus himalayensis)". Animal Biology. 60 (4): 449–465. CiteSeerX doi:10.1163/157075610X523314.
Hartert, E (1921). Die Vogel der palaarktischen Fauna. Volume 3. R. Friedlander & Sohn, Berlin. pp. 1901–1903.
Potapov, R.L. (1993). "New subspecies of the Himalayan Snowcock, Tetraogallus himalayensis sauricus, subsp. nova". Russian Journal of Ornithology. 2 (1): 3–5.
Bianchi V (1899). "Uebersicht der Arten der Gattung Tetraogallus Gray". Journal of Ornithology (in German). 47 (4): 421–434. doi:10.1007/BF02208465. S2CID 34044479.
Koelz, Walter (1951). "Four new subspecies of birds from southwestern Asia". American Museum Novitates (1510). hdl:2246/3970.
Marien, Daniel (1951). "Notes on some pheasants from southwestern Asia, with remarks on molt". American Museum Novitates (1518): 1–25. hdl:2246/3909.
Ali, S; S D Ripley (1980). Handbook of the birds of India and Pakistan. Vol. 2 (2nd ed.). Oxford University Press. pp. 13–16. ISBN 0-19-562063-1.
Stiver SJ (1984). "The Himalayan Snowcock – Nevada's newest upland game" (PDF). Cal-Neva Wildlife Transactions. 26: 55–58.
Ogilvie-Grant, WR (1896). A hand-book to the Game-birds. Volume 1. Edward Lloyd, London. pp. 86–89.
Mayers, James (1985). "Studies of the ecology of Himalayan Snowcock (Tetraogallus himalayensis) in Hunza". World Pheasant Association Journal. 10: 72–86.
Jerdon, TC (1864). The Game birds and Wild fowl of India. Military Orphan Press. pp. 62–63.
Bland, JD; Temple SA (1990). "Effects of predation-risk on habitat use by Himalayan Snowcocks". Oecologia. 82 (2): 187–191. Bibcode:1990Oecol..82..187B. doi:10.1007/BF00323534. PMID 28312664. S2CID 25192814.
Abbott, U.; G. Christensen (1971). "Hatching and rearing the Himalayan snow partridge in captivity". Journal of Wildlife Management. 35 (2): 301–306. doi:10.2307/3799604. JSTOR 3799604.
Baker, EC Stuart (1924). "The game birds of India, Burma and Ceylon, part 38". Journal of the Bombay Natural History Society. 30 (1): 1–11.
Lawrence, WR (1895). The valley of Kashmir. Oxford university Press, London. p. 133.
Junker, K; J Boomker (2007). "A check list of the helminths of guineafowls (Numididae) and a host list of these parasites". Onderstepoort Journal of Veterinary Research. 74 (4): 315–337. doi:10.4102/ojvr.v74i4.118. hdl:2263/5124. PMID 18453241.

Gvosdev, I. (1954). "Helminth fauna of Tetraogallus himalayensis Gray, 1842". Zoologicheskii Zhurnal (in Russian). 33: 39–43.

Other sources

Pheasants, Partridges and Grouse by Madge and McGowan, ISBN 0-7136-3966-0
National Geographic Field Guide to the Birds of North America ISBN 0-7922-6877-6
Handbook of the Birds of the World Vol 2, Josep del Hoyo editor, ISBN 84-87334-10-5
"National Audubon Society" The Sibley Guide to Birds, by David Allen Sibley, ISBN 0-679-45122-6

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