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Superregnum: Eukaryota
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Superphylum: Deuterostomia
Phylum: Chordata
Cladus: Craniata
Subphylum: Vertebrata
Infraphylum: Gnathostomata
Superclassis: Tetrapoda
Cladus: Reptiliomorpha
Cladus: Amniota
Classis: Reptilia
Cladus: Eureptilia
Cladus: Romeriida
Subclassis: Diapsida
Cladus: Sauria
Infraclassis: Lepidosauromorpha
Superordo: Lepidosauria
Ordo: Squamata
Subordo: Gekkota
Infraordo: Gekkomorpha

Superfamilia: Eublepharoidea
Familia: Eublepharidae
Genus: Goniurosaurus
Species (23): G. araneus – G. bawanglingensis – G. catbaensis – G. gezhi – G. gollum – G. hainanensis – G. huuliensis – G. kadoorieorum – G. kuroiwae – G. kwanghua – G. kwangsiensis – G. liboensis – G. lichtenfelderi – G. luii – G. orientalis – G. sengokui – G. splendens – G. toyamai – G. varius – G. yamashinae – G. yingdeensis – G. zhelongi – G. zhoui

Goniurosaurus Barbour, 1908: 316

Type species: Goniurosaurus hainanensis Barbour, 1908, by monotypy.


Amamisaurus Börner, 1981: 4 [synonymised by Grismer (1987: 44)]
Type species: Gymnodactylus albofasciatus kuroiwae Namiye, 1912, by original designation.

Primary references

Barbour, T. 1908. Some new reptiles and amphibians. Bulletin of the Museum of Comparative Zoology 51(12): 315–325. BHL
Börner, A.-R. 1981. The genera of Asian eublepharine geckos and a hypothesis on their phylogeny. Miscellaneous Articles in Saurology 9(1): 1–14. Reference page.
Grismer, L.L. 1987. Evidence for the resurrection of Goniurosaurus Barbour (Reptilia: Eublepharidae) with a discussion on geographic variation in Goniurosaurus lichtenfelderi. Acta Herpetologica Sinica 6(1): 43–47. BHL Reference page.

Additional references

Grismer, L.L. 1988. Phylogeny, taxonomy, classification, and biogeography of eublepharid geckos. pp. 369–469 In: Estes, R. & Pregill, G. (eds.). Phylogenetic Relationships of the Lizard Families: Essays Commemorating Charles L. Camp. Stanford University Press: Stanford, California. ISBN 978-0804714358 Reference page.
Grismer, L.L., Ota, H. & Tanaka S. 1994. Phylogeny, classification, and biogeography of Goniurosaurus kuroiwae (Squamata: Eublepharidae) from the Ryukyu Archipelago, Japan, with description of a new subspecies. Zoological Science 11(2): 319–335. BHL Reference page.
Grismer, L.L., Viets, B.E. & Boyle, L.J. 1999. Two New Continental Species of Goniurosaurus (Squamata: Eublepharidae) with a Phylogeny and Evolutionary Classification of the Genus. Journal of Herpetology 33(3): 382–393. DOI: 10.2307/1565635 Reference page.
Orlov, N.L. & Daresvky, I.S. 1999. Description of a New Mainland Species of Goniurosaurus Genus, from the North-Eastern Vietnam. Russian Journal of Herpetology 6(1): 72–78. DOI: 10.30906/1026-2296-1999-6-1-72-78 [nonfunctional] Broken access Reference page.
Ota, H., Honda, M., Kobayashi, M., Sengoku, S. & Hikida, T. 1999. Phylogenetic Relationships of Eublepharid Geckos (Reptilia: Squamata): A Molecular Approach. Zoological Science 16(4): 659–666. DOI: 10.2108/zsj.16.659 Open access Reference page.
Grismer, L.L. 2000. Goniurosaurus murphyi Orlov and Darevsky: A Junior Synonym of Goniurosaurus lichtenfelderi Mocquard. Journal of Herpetology 34(3): 486–488. DOI: 10.2307/1565379 Paywall Reference page.
Grismer, L.L., Haitao, S., Orlov, N.L. & Ananjeva, N.B. 2002. A New Species of Goniurosaurus (Squamata: Eublepharidae) from Hainan Island, China. Journal of Herpetology 36(2): 217–224. DOI: 10.2307/1565994 Paywall Reference page.
Orlov, N.L., Ryabov, S.A., Nguyen, T.T., Nguyen, T.Q. & Ho, C.T. 2008. A New Species of Goniurosaurus (Sauria: Gekkota: Eublepharidae) from North Vietnam. Russian Journal of Herpetology 15(3): 229–244. DOI: 10.30906/1026-2296-2008-15-3-229-244 Paywall Reference page.
Ziegler, T., Truong, N.Q., Schmitz, A., Stenke, R. & Rösler, H. 2008. A new species of Goniurosaurus from Cat Ba Island, Hai Phong, northern Vietnam (Squamata: Eublepharidae). Zootaxa 1771(1): 16–30. DOI: 10.11646/zootaxa.1771.1.2 Paywall. PDF from Researchgate. Reference page.
Wang, Y.-Y., Yang, J.-H. & Cui, R.-F. 2010. A New Species of Goniurosaurus (Squamata: Eublepharidae) from Yingde, Guangdong Province, China. Herpetologica 66(2): 229–240. DOI: 10.1655/09-046R2.1 Paywall Reference page.
Wang, Y.-Y., Yang, J.-H. & Grismer, L.L. 2013. A New Species of Goniurosaurus (Squamata: Eublepharidae) from Libo, Guizhou Province, China. Herpetologica 69(2): 214–226. DOI: 10.1655/HERPETOLOGICA-D-12-00084 Reference page.
Honda, M., Kurita, T., Toda, M. & Ota, H. 2014. Phylogenetic Relationships, Genetic Divergence, Historical Biogeography and Conservation of an Endangered Gecko, Goniurosaurus kuroiwae (Squamata: Eublepharidae), from the Central Ryukyus, Japan. Zoological Science 31(5): 309–320. DOI: 10.2108/zs130201 Open accessReference page.
Wang, Y.-Y., Jin, M.-J., Li, Y.-L. & Grismer, L.L. 2014. Description of a New Species of Goniurosaurus (Squamata: Eublepharidae) from the Guangdong Province, China, Based on Molecular and Morphological Data. Herpetologica 70(3): 309–322. DOI: 10.1655/09-046R2.1Reference page.
Yang, J-H. & Chan, B-P. 2015. Two new species of the genus Goniurosaurus (Squamata: Sauria: Eublepharidae) from southern China. Zootaxa 3980(1): 67–80. DOI: 10.11646/zootaxa.3980.1.4 Preview (PDF). Full article (PDF). Reference page.
Honda, M. & Ota, H. 2017. On the Live Coloration and Partial Mitochondrial DNA Sequences in the Topotypic Population of Goniurosaurus kuroiwae orientalis (Squamata: Eublepharidae), with Description of a New Subspecies from Tokashikijima Island, Ryukyu Archipelago, Japan. Asian Herpetological Research 8(2): 96–107. DOI: 10.16373/j.cnki.ahr.170003 Reference page.
Kurita, T., Honda, M. & Toda, M. 2018. Species delimitation and biogeography of the Ryukyu ground geckos, Goniurosaurus kuroiwae ssp. (Squamata: Eublepharidae), by use of mitochondrial and nuclear DNA analyses. Journal of Zoological Systematics and Evolutionary Research 56(2): 209–222. DOI: 10.1111/jzs.12198 Reference page.
Zhou, R-B., Wang, N., Chen, B. & Liang, B. 2018. Morphological evidence uncovers a new species of Goniurosaurus (Squamata: Eublepharidae) from the Hainan Island, China. Zootaxa 4369(2): 281–291. DOI: 10.11646/zootaxa.4369.2.8 Paywall Reference page.
Qi, S., Grismer, L.L., Lyu, Z.-T., Zhang, L., Li, P.-P. & Wang, Y.-Y. A definition of the Goniurosaurus yingdeensis group (Squamata, Eublepharidae) with the description of a new species. ZooKeys 986: 127–155. DOI: 10.3897/zookeys.986.47989 Open access Reference page.
Qi, S., Wang, J., Grismer, L.L., Chen, H.-H., Lyu, Z.-T. & Wang, Y.-Y. 2020. The Stoor Hobbit of Guangdong: Goniurosaurus gollum sp. nov., a cave-dwelling Leopard Gecko (Squamata, Eublepharidae) from South China. ZooKeys 991: 137–153. DOI: 10.3897/zookeys.991.54935 Open access Reference page.
Zhu, X.-Y., Shen, C.-Z., Liu, Y.-F., Chen, L., Li, Z. & He, Z.-Q. 2020. A new species of Goniurosaurus from Hainan Island, China based on molecular and morphological data (Squamata: Sauria: Eublepharidae). Zootaxa 4772(2): 349–360. DOI: 10.11646/zootaxa.4772.2.6 Paywall Reference page.
Zhu, X.-Y., Chen, G.-Y., Román-Palacios, C., Li, Z. & He, Z.-Q. 2020. Goniurosaurus gezhi sp. nov., a new gecko species from Guangxi, China (Squamata: Eublepharidae). Zootaxa 4852(2): 211–222. DOI: 10.11646/zootaxa.4852.2.6 Paywall Reference page.


Uetz, P. & Hallermann, J. 2021. Goniurosaurus . The Reptile Database. Accessed on 17 September 2018.
Goniurosaurus – Taxon details on Interim Register of Marine and Non-marine Genera (IRMNG).
Goniurosaurus – Taxon details on Integrated Taxonomic Information System (ITIS).

Vernacular names
English: East Asian Leopard Geckos

Goniurosaurus is a genus of geckos, containing 25 species. Members are known by various names including ground geckos, tiger geckos, leopard geckos, and cave geckos Members of this genus are found in China, Japan, and Vietnam. For this reason they are known commonly as Asian geckos. They belong to the family Eublepharidae.


Geckos of the genus Goniurosaurus mainly feed on small insects.

The following 25 species are recognized as being valid.[1][2]

They can be divided into 4 species groups.

luii group:

Goniurosaurus araneus Grismer et al., 1999 (Vietnamese leopard gecko)
Goniurosaurus bawanglingensis Grismer et al., 2002
Goniurosaurus catbaensis Ziegler, Truong, Schmitz, Stenke & Rösler, 2008 (Cat Ba leopard gecko)
Goniurosaurus chengzheng Zhu, Liu, Bai, Roman-Palacios, Li, & He, 2021 (Chengzheng cave gecko)
Goniurosaurus gezhi Zhu, He & Li, 2020[3]
Goniurosaurus huuliensis Orlov et al., 2008
Goniurosaurus luii Grismer et al., 1999
Goniurosaurus liboensis Wang et al., 2013
Goniurosaurus kadoorieorum Yang & Chan, 2015 (Kadoories's cave gecko)
Goniurosaurus kwangsiensis Yang & Chan, 2015 (Guangxi cave gecko)

lichtenfelderi group:

Goniurosaurus hainanensis Barbour, 1908
Goniurosaurus kwanghua Zhu & He, 2020
Goniurosaurus lichtenfelderi (Mocquard, 1897) (Lichtenfelder's gecko)
Goniurosaurus sinensis Zhou, Peng, Hou, & Yuan, 2019
Goniurosaurus zhoui Zhou, Wang, Chen, & Liang, 2018 (Zhou's leopard gecko)

kuroiwae group:

Goniurosaurus kuroiwae (Namiye, 1912) (Kuroiwa's ground gecko)
Goniurosaurus orientalis (Maki, 1931) (Spotted ground gecko)
Goniurosaurus sengokui (Honda & Ota, 2017) (Sengoku's gecko)
Goniurosaurus splendens (Nakamura & Uéno, 1959) (Banded ground gecko)
Goniurosaurus toyamai Grismer et al., 1994 (Iheyajima leopard gecko)
Goniurosaurus yamashinae (Okada, 1936) (Yamashina's ground gecko)

yingdeensis group:

Goniurosaurus yingdeensis Wang et al., 2010 (Yingde leopard gecko)
Goniurosaurus varius Qi et al., 2020
Goniurosaurus zhelongi Wang et al., 2014 (Zhe-long's leopard gecko)


Goniurosaurus gollum Qi et al., 2020 (Gollum leopard gecko)


"Goniurosaurus ". The Reptile Database.
Qi, Shuo; Grismer, L. Lee; Lyu, Zhi-Tong; Zhang, Liang; Li, Pi-Peng; Wang, Ying-Yong (2020-11-05). "A definition of the Goniurosaurus yingdeensis group (Squamata, Eublepharidae) with the description of a new species". ZooKeys (986): 127–155. doi:10.3897/zookeys.986.47989. ISSN 1313-2970. PMC 7661482. PMID 33223883.

Zhu, Xiao-Yu; Chen, Guang-Yu; Román-Palacios, Cristian; Li, Zheng; He, Zhu-Qing (2020-09-15). "Goniurosaurus gezhi sp. nov., a new gecko species from Guangxi, China (Squamata: Eublepharidae)". Zootaxa. 4852 (2): 211–222. doi:10.11646/zootaxa.4852.2.6. ISSN 1175-5334. PMID 33056426. S2CID 222821642.

Further reading

Barbour T (1908). "Some New Reptiles and Amphibians". Bull. Mus. Comp. Zool., Harvard College 51 (12): 315–325. (Goniurosaurus, new genus, p. 316).

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