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Superregnum: Eukaryota
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Superphylum: Deuterostomia
Phylum: Chordata
Cladus: Craniata
Subphylum: Vertebrata
Infraphylum: Gnathostomata
Superclassis: Tetrapoda
Cladus: Reptiliomorpha
Cladus: Amniota
Classis: Reptilia
Cladus: Eureptilia
Cladus: Romeriida
Subclassis: Diapsida
Cladus: Sauria
Infraclassis: Lepidosauromorpha
Superordo: Lepidosauria
Ordo: Squamata
Cladus: Unidentata Episquamata Toxicofera
Subordo: Iguania
Infraordo: Acrodonta

Familia: Agamidae
Subfamilia: Draconinae
Genus: Gonocephalus
Species (16): G. abbotti – G. bellii – G. beyschlagi – G. bornensis – G. chamaeleontinus – G. doriae – G. grandis – G. interruptus – G. klossi – G. kuhlii – G. lacunosus – G. liogaster – G. megalepis – G. mjoebergi – G. semperi – G. sophiae

Gonocephalus Kaup, 1825: 590

Type species: Gonyocephalus tigrinus Wagler, 1830 [=Lophyrus tigrinus Duméril & Bibron, 1837 fide Wermuth (1967: 55)], by subsequent designation by Fitzinger (1843: 15).

Goniocephalus Kaup, 1827: 614 [unjustified emendation]
Gonyocephalus Wagler, 1830: 150 [incorrect subsequent spelling]

Primary references

Kaup, J.J. 1825. Einige Bemerkungen zu Merrems Handbuch. Isis von Oken 16: 589–593. BHL

Additional references

Denzer, W., Manthey, U., Mahlow, K. & Böhme, W. 2015. The systematic status of Gonocephalus robinsonii Boulenger, 1908 (Squamata: Agamidae: Draconinae). Zootaxa 4039(1): 129–144. DOI: 10.11646/zootaxa.4039.1.5. Preview (PDF) Reference page.


Uetz, P. & Hallermann, J. 2021. Gonocephalus . The Reptile Database. Accessed on 28 February 2019.

Vernacular names
Deutsch: Winkelkopfagamen
日本語: モリドラゴン属

Gonocephalus is a genus of agamid lizards endemic to southeast Asia.[4]


Species from Melanesia and Australia which were formerly included in Gonocephalus are now placed in the genus Hypsilurus.[4]

As of 2013, Gonocephalus contains the following 17 species:[3]

Gonocephalus abbotti Cochran, 1922 – Abbott's crested lizard
Gonocephalus bellii (A.M.C. Duméril & Bibron, 1837) – Bell's anglehead lizard, Bell's forest dragon
Gonocephalus beyschlagi (Boettger, 1892) – Sumatra forest dragon
Gonocephalus bornensis (Schlegel, 1848) – Borneo anglehead lizard, Borneo forest dragon
Gonocephalus chamaeleontinus (Laurenti, 1768) – chameleon forest dragon, chameleon anglehead lizard
Gonocephalus doriae (W. Peters, 1871) – Doria's angle-headed lizard
Gonocephalus grandis (Gray, 1845) – giant forest dragon, great anglehead lizard
Gonocephalus interruptus (Boulenger, 1885) – Boulenger's forest dragon, Mindoro anglehead
Gonocephalus klossi (Boulenger, 1920) – Kloss's forest dragon
Gonocephalus kuhlii (Schlegel, 1848)
Gonocephalus lacunosus Manthey & Denzer, 1991 – Manthey's forest dragon
Gonocephalus liogaster (Günther, 1872) – tropical forest dragon, blue-eyed anglehead lizard, orange-ringed anglehead lizard,
Gonocephalus megalepis (Bleeker, 1860) – Bleeker's forest dragon
Gonocephalus mjobergi M.A. Smith, 1925
Gonocephalus pyrius Harvey, Rech, Riyanto, Kurniawan & E.N. Smith, 2021 – angle-headed dragon
Gonocephalus semperi (W. Peters, 1867) – Mindoro forest dragon
Gonocephalus sophiae (Gray, 1845) – Negros forest dragon

Nota bene: A binomial authority in parentheses indicates that the species was originally described in a genus other than Gonocephalus.

Kaup JJ ("Caup [sic]") (1825). "Einige Bemerkungen zu Merrems Handbuch ". Isis von Oken 16: 589-593. (Gonocephalus, new genus, p. 590). (in German).
Boulenger GA (1885). Catalogue of the Lizards in the British Museum (Natural History). Second Edition. Volume II. ... Agamidæ. London: Trustees of the British Museum (Natural History). (Taylor and Francis, printers). xiii + 497 pp. + Plates I-XXIV. (Genus Gonyocephalus [sic], p. 282).
Gonocephalus at the Reptile Database. Accessed 5 September 2013.

Honda M, Ota H, Sengoku S, Yong HS, Hikida T (2002). "Molecular Evaluation of Phylogenetic Significances in the Highly Divergent Karyotypes of the Genus Gonocephalus (Reptilia: Agamidae) from Tropical Asia". Zoological Science. 19 (1): 129–133. doi:10.2108/zsj.19.129. hdl:2241/104127. PMID 12025399. S2CID 38549668.

Wikispecies has information related to Gonocephalus.
Further reading

Das I (2006). A Photographic Guide to Snakes and other Reptiles of Borneo. Sanibel Island, Florida: Ralph Curtis Books. 144 pp. ISBN 0-88359-061-1. (Gonocephalus species, pp. 81–84).
Smith MA (1935). The Fauna of British India, Including Ceylon and Burma. Reptilia and Amphibia. Vol. II.—Sauria. London: Secretary of State for India in Council. (Taylor and Francis, printers). xiii + 440 pp. + Plate I + 2 maps. (Genus Goniocephalus [sic], p. 157).

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